Can Plants Communicate
Can Plants Communicate
Can Plants Communicate
Introduction
All living organisms begin in the same form: as a single cell. That cell
will divide and the resulting cells will continue dividing and differentiate
into cells with various roles to carry out within the organism. This is life
and plants are no different. Plant growth can be determinate or
indeterminate, meaning some plants will have a cycle of growth then a
cessation of growth, breakdown of tissues and then death (think of a
radish plant or a tomato plant) while others (think of a giant cedar tree)
will grow and remain active for hundreds of years. A tomato plant is fairly
predictable and is said to have determinate growth, while the cedar tree
has indeterminate growing potential. Development refers to the growth
and differentiation of cells into tissues, organs and organ systems. This
again all begins with a single cell.
Genetic information directs the synthesis and development of
enzymes which are critical in all metabolic process within the plant. Most
enzymes are proteins in some form or another, are produced in very
minute quantities and are produced on sitemeaning they are not
transported from one part of the organism to another. Genetic information
also regulates the production of hormones, which will be addressed shortly.
The major difference is that hormones are transported from one part of the
plant to another as needed. Vitamins vital in the activation of enzymes
and are produced in the cytoplasm and membranes of plant cells. Animals
and humans utilize plants in order to provide some vitamin resources. In
general, hormone and vitamin effects are similar and are difficult to
distinguish in plants, and both are referred to in general as plant growth
regulators.Occasionally we hear or read of experiments often associated
with school science fairs-that suggest plants respond in some positive way
to good music or soothing talk ; conversely, some plants are said to grow
poorly when exposed to loud rock music or to being harshly yelled at.
There is currently much debate about global warming and the
potential effects on life as we know it. Are those who proclaim that global
CHAPTER II
DISCUSSION
What are Plant Hormones ?
Plant hormones affect how plants grow. Unlike animal hormones, plant hormones are
organic molecules. They aid in plant growth and survival. Cellular events like division and
cell morphology are driven by hormone levels. In plants, meristematic tissue is composed of
cells that are actively growing. This is where one is expected to find a relatively large
concentration of plant hormones. Also, one will find higher hormone levels in root tips.
The major known types of plant hormones are auxins, gibberelins, cytokinins, abscis
acid, and ethylen.
Regulating Growth: Plant Hormones
Plant cells are in constant chemical communication with one another and with their
environment. They recognize and respond to stimuli of many kinds, using a system of
chemical messengers that receive and transmit the stimuli via ordinary body cells (unlike the
highly specialized cells of animal nervous systems). Control of the plant system apparently
resides in the genes of each cell, which are turned on and off by the chemical messages they
receive. The response may be stimulatory (initiating cellular division and enlargement, for
example) or inhibitory (such as stopping a metabolic process)
The chemical messengers are hormones, organic substances manufactured in small
amounts in one tissue and usually transported to another where they initiate a response. (A
few act in the tissues where they are produced.) The hormone molecule itself carries little
information and produces a reaction only when it binds to appropriate receptor molecules at
the response site.
Plants, in comparison to animals, have both fewer hormones and fewer kinds of
responses. Plant hormones, however, usually act in combination, thus producing more varied
responses than if acting individually. The same hormone also can produce different effects
when acting in different tissues or in different concentrations in the same tissue. The
developmental stage of the plant additionally determines what effects the hormone activates.
Growth and development depend upon a successful coordination of the activities of
hormones, not just the presence or absence of individual ones
There must be good recognition and strong binding between the hormone
and the responding molecules.
There are two generally accepted mechanisms by which hormones will act.
1. The first type deals with a steroid hormone. In this type the hormone
can pass through the plasma membrane into the cytoplasm. Here it
binds with its receptor molecule to form a hormone-receptor
complex. From this point, the complex may dissociate (If there is not
tight binding) or it may enter the nucleus and affect mRNA
synthesis. The effect of the hormone on mRNA synthesis ultimately
results in the physiological response(Arteca,1996 Moran et.al., 1994;
Wolfe, 1993)
are not ripe, shoots, leaves and roots (Rismunandar, 1988). GA biosynthesis involves 3
chemical metabolites, ie mevalonic acid that acts as a precursor for the formation of
isoprene, which is the base of the carbon-19 and carbon-20 framework giban, kaurena
formed from isoprene, GA formed from kaurena (Leopold and Kriedemann, 1975 in
Gardner, et al ., 1991).
Gibberellins work synergistically with auxin, cytokinins and perhaps some
other substances (synergism) to influence the peak dormancy, growth cambium,
geotropisme, abscission and partenokarpi (due to the activity of auxin and gibberellin),
effectively increasing fruit set, thus stimulating the growth of inter-books are not
stunted plants , the release of a-amylase for starch hydrolysis and germination
(Gardner, et al., 1991). Giberilin reacts in the cells surrounding the endosperm which
led to the formation of a number of specific hydrolytic enzymes (such as amylase and
protease) that digests starch and protein endosperm dengam thus making supplies
sugars and amino acids for cell growth (Kimball, 1983). He also explained that the
amino acids provided by the action of the protease enzyme is precurson formation of
another type of growth hormone, as triptopan which is an early form of auxin.
According Kusumo (1989), that gibberellins play a role in cell division and
supports the formation of RNA resulting in protein synthesis. Cell division is
stimulated by active amylase hydrolyzes starch into sugar is reduced so that the
increased sugar concentration also increased as a result of the osmotic pressure.
Increased osmotic pressure inside the cell causing water to easily enter the cells, so it
can be triggers all physiological processes in plant cells.
Gibberellins function is to stimulate cell division and stimulate the activity of
amylase and proteinase enzymes that play a role in germination. Gibberellins also
stimulates the formation of buds, remove seed dormancy, and stimulate fruit growth
by parthenogenesis.
3) Cytokines
Cytokines in accordance with its name derived from the cytokines are growth
hormones that affect cell division. According to Kimball (1983), cytokines when
reacted with Auxin, strongly stimulate mitosis in tissue merestematik, a real explosion
of RNA synthesis occurs when plant cells or isolated nuclei treated with cytokines.
Furthermore, according to Wereing and Philips (1981), in the metabolic processes of
cytokines thought to have an important role in protein synthesis, the process of
translation.
Cytokines can be found on the network are split as follows: At the root,
embryo and fruit, to move from the roots to other organs. Cytokinins were discovered
first is kinetin. Contained cytokinin is zeatin in Zea mays. The function of cytokines is
Stimulate cell division, stimulate the formation of buds on the stem and on the
callus, inhibits the effects of apical dominance, and accelerate the growth of elongated
4) Abscisic acid
Not all hormones serve to stimulate growth, because there are also inhibits
growth, abscisic acid. Abscisic acid (ABA), as growth inhibitors (Inhibitor / retardant)
when plants experience stress, is used to mengompakkan fitohormon stem growth so
that plants look very good. Abscisic acid function is to inhibit cell division and
elongation, delayed growth or dormancy, stimulate closure leaves the mouth dry
season, and helped the decay of leaves in the dry season
Place it generates: Leaf; stems, roots, fruits green.
5) Ethylene
Ethene or ethylene is the simplest alkene compounds consisting of four
hydrogen atoms and two carbon atoms connected by a double bond. Because of this
bond, ethene also called unsaturated hydrocarbons or olefins.
At room temperature, the molecules can not rotate the ethene double bond so
that all constituent atoms are on the same plane. The angle formed by two carbonhydrogen bonds in the molecule is 117 , very close to 120 are estimated based on
ideal sp2 hybridization. Ethylene function is to encourage ripening; give the opposite
effect with some influence auxin; encourage or inhibit the growth and development of
roots, leaves, stems and flowers.
Place generates: Ripe fruit, a book on the stem, leaf aging.
6) Brassinolide (brassinosteroid group)
Brassinolide is similar fitohormon steroids in animals and have a response
similar to gibberellin. Several functions brassinolide are as follows: increase the rate
of plant cell elongation, inhibit leaf aging (senescence), resulting in a bend on the
leaves of grasses, inhibiting the leaf drop, inhibiting the growth of plant roots, shoots
of plants increases resistance to environmental stress, stimulates cell renewal in shoots
of plants, stimulate the growth of plant shoots, stimulating plant xylem differentiation,
inhibit the growth of shoots at deficient (deficient) and endogenous carbohydrate air.
Brassinolide synthesized from acetyl CoA via the acid mevalonik.
7) Polyamines
Polyamines have a major role in the most fundamental genetic processes such
as DNA synthesis and genetic expression. Spermine and spermidine bind to phosphate
chains of nucleic acids. This interaction kebanyakkan based on electrostatic interaction
between the positively charged ions of ammonium groups of the polyamine and the
negative charge of the phosphate.
Polyamine is the key to cell migration, proliferation and differentiation in
plants and animals. Level metabolic precursor of polyamine and amino acids is very
fields. Indolebutyric acid (IBA) and naphthalene acetic acid (NAA) are commonly used to
promote root development in stem cuttings. Treating apple and pear trees with NAA just after
their full bloom will cause a thinning of up to 80% of the developing fruit, a situation that is
desirable because fewer large fruit have a greater market value than many small fruit. Later in
fruit
development,
NAA,
2,4-dichlorophenoxyacetic
acid
(2,4-D),
and
2,4,5-
trichlorophenoxypropionic acid (2,4,5-TP) may be used to delay fruit drop, extending the
harvest season. Many farmers, as well as many homeowners, will use 2,4-D as an herbicide to
kill broadleaf weeds in their fields or lawns.
Of the more than 90 forms of gibberellin that have been identified in higher plants and
in the fungus Gibberella, only the forms GA4 and GA7 have been found to have commercial
value. These gibberellins are used extensively to increase the size and quality of seedless
grapes. They are used to delay the ripening of lemons so that these can be harvested during
the months with the greatest demand. Application of gibberellins to artichokes promotes the
production of flower buds and allows for earlier harvesting dates. These gibberellins are also
used to increase the yields of malt from barley and sucrose from sugarcane.
The cytokinin-like compounds, benzyladenine and tetrapyranylbenzyladenine, are
used to promote lateral branching in white pine and carnations, respectively. A combination of
benzyladenine, GA4, and GA7, called Promalin, is used to promote the elongation of the
Delicious apple varieties so that they have the elongated shape expected by the public, rather
than a rounder shape. Promalin is also applied to young fruit trees to increase lateral
branching, producing a tree with the desired shape for mechanical harvesting.
Several treatments or compounds that increase ethylene concentrations in or around
the plants have found commercial uses. Among the oldest of these treatments is the burning of
fires or running of gas engines adjacent to fields of pineapple and mango, helping to
synchronize flowering in these plants. The incomplete combustion of fuels in both of the
cases adds ethylene to the environment to achieve the desired effect. A compound commonly
applied
to Hevea
brasiliensis (the
tree
that
produces
natural
rubber)
is
2-
bind to receptor molecules, usually located in the plasma membrane, and thereby
trigger a response. Hormones appear to be released into general circulation and are not
carried specifically to the target. Many regions that are not target regions are exposed
to the hormone but do not respond because they do not have the proper receptor
molecules. In some instances, a plant hormone acts directly on the cells that produce
it.
At one time, plant hormones were believed to carry in their structure much of
the information necessary for the response. We now know that plant hormones are
quite simple in structure. The receptor cell and its nucleus contain almost all of the
information necessary for proper response, and hormones serve only to activate the
response.
Can Plants Move Rapidly ?
movements result from varying growth rates in diferent parts of an organ. They are
mainly related to young parts of a plant and as a rule are quite slow, usually taking at least 2
hours to become apparent, although the plant may have begun microscopic changes within
minutes of reiving a stimulus.
Movements Resulting Primarily from Internal Stimuli
Nutations, Charles Darwin once attached a tiny sliver of glass to the tip of a plant growing in
a pot. The he suspended a piece of paper blackened with carbon over the tip, and as the plant
grew. He raised the paper just enough to allow the tip to touch the paper without hurting the
plant. He found that the growing point traced a spiral pattern in the blackened paper. We know
now that such nutations (also referred to as spiraling movements) are common to many plants.
Nodding Movements, members of the Legume Family (Fabaceae), such as garden beans,
whose ethylene production upon germinating causes the formation of a thickened crook in the
hypocotyl, exhibit a slow oscillating movement (i,e.. the bent hypocotyl nods from side to
side like an upside down pendulum) as the seedling pushes up through the soil. This nodding
movement apparently facilitates the progress of the growing plant tip through the soil.
Twining Movements, although twining movements are mostly stimulated internally, external
forces, such as gravity and contact, may also play a role. These movements occur when ncells
in the stems of climbing plants, such as morning glory, elongate to differing extents, causing
visible spiraling in growth (in contrast with the spiraling movements previosly mentioned,
which are not visible to the eye). Tendtril twining, which is initiated by contact, results from
an elongation of cells on one side, followes by differences in growth rates. Some tendtrils are
stimulated to coil by auxin, while others are stimulated by ethylene.
Contraction Movements, we noted that the bulbs of a number of dicots and monocots have
contractile roots, which pull them deeper into the ground. In lilies, for example, seeds
germinating at the surface ultimately produce bulbs that end up 10 to 15 centimeters (4 to 6
inches) below ground level because of the activities of contractileroots. There is some
evidence that temperature fluctuations at the surface determine how long the contracting will
continue. When the bulb gets deep enough that the differences between daytime and nightime
temperatures are slight, the contractions cease.
Nastic movements, when flattened plant organs, such as leaves or flower petals, first
expandfrom buds, they characteristically alternative in bending down and then upas the cells
in the upper and lower parts of the leaf alternate in enlarging faster than those in the opposite
parts. Such nondirectional movements (i.e.., movements that do not result in an organ being
oriented toward or away from the direction of a stimulus) are called nastic. Nastic movements
may involve differential growth or turgor changes in special cells. Epinasty is the permanent
downward bending of organ, often the petiole of a leaf, in response to either an unequal flow
of auxin through the petiole or to ethylene. Nastic movements that involve changes in
turgorpressure include sleep movements and contact movements, which are discussed later.
Movements Resulting from External Stimuli
Permanent movements resulting from external stimuli coming from one direction are
commonly referred to as tropisms. Tropic movements can be divided into three phases:
1. In the initial perception phase, the organ receives a greater stimulus on one side.
2. Then, transduction occurs, during which one or more hormones becomes unevenly
distributed across the organ.
3. Finally, asymmetric growth occurs as a result of the uneven distribution of the
hormone causing greater cell elongation on one side.
Phototropism. The main shoots of most plants growing in the open tend to develop vertically,
although the branches often grow horizontally. If a box is placed over a plant growing
vertically and a hole is cut to admit light from one side. The tip of the plant will begin to bend
toward the light within a few hours. If the box is later removed, a compensating bend
develops, causing the tip to grow vertically again. Such a growth movement toward light is
called a positive phototropism. A similiar bending away from light is called a negative
phototropism. The shoots tips of most plants are positively phototropic, while roots are either
insensitive to light or negatively phototropic.
Gravitropism. Growth responses to the stimulus of gravity are called gravitropisms. The
primary roots of plants are positively gravitropic, while shoots forming the main axis of plants
are negatively gravitropic. It was postulated that plant organs perceive gravity through the
movement of amyloplasts containing large starch grains located in special cells of the root
cap. The amyloplasts are also found in coleoptile tips and in the endodermis. When a potted
plant is placed on its side, the starch containing amyloplasts will, within a few minutes, begin
to float ao ruble down until they come to rest on the side of the cells closest to the gravity
stimulus. In roots, the cells on the side opposite the stimulus begin elongating within 10
seconds to an hour or two, bringing about a downward bend, while the opposite occurs in
stems.
Other Tropisms. A plant or plant part response to contact with a solid object is called a
thigmotropism. One of the most common thigmotropic responses is seen in the coiling of
tendrils and in the twining of climbing plant stems. Such responses can be relatively rapid,
with some tendrils wrapping around a support two or more times within an hour. The coiling
results from cells in contact becoming slightly shorter while those on the opposite side
elongate.
CHAPTER III
CONCLUSION
Plant hormones is a collection of organic compounds not nutrient (nutrients)both naturally
occuring and man-made, which in very small levels could encourage, inhibit, or alter the
growth, development, and movement (taksis) plants. Level of small is in the range of one
milimoles per liter one micromol per liter.
Types of Hormones In Plant And Their Function
1. Auxin is to assist in the process of accelerating growth, the growth of
both root and stem growth, accelerate germination, helps in the
process of fruit ripening sel.mempercepat division, reducing the
number of seeds in the fruit and aging, and abortion
2. Giberelin is to Stimulate cell division cambium,Stimulate early flowering
prematurely, Stimulate the formation of seedless fruit, Stimulate plaant grows very
quickly so as to have a giant size
3. Cytokinin to Stimulate the process of cell division, Delaying defoliaton, flower and
fruit, Affect the growth of shoots and roots, Enhace resistance to adverse effects such
as loe temperature, viral invections, weed killers and radiation, Inhibit (hold)
yellowing of the leaves by making the proteins and chlorophyll in the leaves of a
balanced
4. Ethylene gas to Help break dormancy in plants, Support the ripening fruit, Support the
abcission on the leaves, Supporting the flowering process, Inhibit root elongation in
some species of plants and can stimulate stem elongation, Stimulate germinate,
Support the formation of root hairs
5. Kalin , Rhizokalin to affecting root formation, Kaulokalin to affects the formation of
stem, Filokalin to affects leaf formation, Antokalin to affects the formation of flowers
6. Abscisic acid (ABA) to Inhibit seed germination, Affect the flowering plants, Extend
the dormancy of tubers, Affect bud dormancy plants to perform
7. Traumalin Acid / Acid traumalat Fix injury in plants (restitution process /
regeneration)
8. Brassinolide (brassinosteroid group) to Increase the rate of plant cell elongation,
Resulting in a bend on the leaves of grasses, Inhibiting the leaf drop, Inhibiting the
growth of plant roots
REFERENCES
Stek
Pucuk
Meranti
Putih
(Shorea
montigena).
Jurusan
Benih
Sukun.
Pusat
Penelitian
dan
Pengembangan
Sutrisno.
2006.
Sukun
Sebagai
Cadangan
Pangan