Harley has an identity crisis that sends her into a strange brain freeze. Dr. Psycho leads the team into Harley's mind to find the source of the problem.
Harley recruits her friend and mentor Queen of Fables to help her steal a force field from STAR Labs, to steal a weather-changing machine, so she can impress the Legion of Doom. Meanwhile, Poison Ivy dates Kite Man.
Ivy and the crew find themselves abandoned when Harley pursues her dream after the Legion makes her a member - and Joker woos her. But when he uses her to escape Batman, Harley realizes that she's made a mistake...but it may be too late to fix things with her friends.
Harley discovers that she can go home again - but it's very dangerous. As Harley reconciles with her father, Ivy tries to escape from the factory and Frank drives to the rescue.
Ivy is dead, Harley's crew is captured, and Batman is his captive. Joker has everything he's ever wanted - but is it enough without Harley at his side?