Ulmus alata

Ulmus alata
Leaves of U. alata
Conservation status
Least Concern  (IUCN 3.1)[1]
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Rosids
Order:	Rosales
Family:	Ulmaceae
Genus:	Ulmus
Subgenus:	U. subg. Oreoptelea
Section:	U. sect. Chaetoptelea
Species:	U. alata
Binomial name
Ulmus alata Michx.
Natural range
Synonyms
Ulmus pumila Walter

Ulmus alata, the winged elm or wahoo, is a small- to medium-sized deciduous tree endemic to the woodlands of the southeastern and south-central United States. The species is tolerant of a wide range of soils, and of ponding, but is the least shade-tolerant of the North American elms. Its growth rate is often very slow, the trunk increasing in diameter by less than 5 mm (3⁄16 in) per year. The tree is occasionally considered a nuisance as it readily invades old fields, forest clearings, and rangelands, proving particularly difficult to eradicate with herbicides.^[2]

As its common and scientific^[3] names imply, winged elm is most easily recognized by the very broad, thin pair of corky wings that form along the branchlets after a couple of years. The tree generally grows to a maximum height and breadth of about 13 m × 13 m (43 ft × 43 ft), although on fertile alluvial soils such as those of the Mississippi River Delta, some specimens have reached double this height (see 'Notable trees' below). The crown can be either rounded or pyramidal; the branches are pendulous.^[4] The leaves are comparatively small for the genus, less than 6.5 cm (2+1⁄2 in) long and less than 2.0 cm (3⁄4 in) broad, oblong-lanceolate to narrowly elliptic, thin in texture, and smooth above with serrate or doubly serrate margins. The leaves turn bright yellow in Autumn. The wind-pollinated perfect apetalous flowers are borne on long pedicels in March and April before the leaves appear. The reddish samaras are also relatively small, less than 8 mm (5⁄16 in) long, narrowly elliptic with two long incurving stigmas at the tip,^[5] and usually disperse before the end of April.^[6][7]

• 
  Young Ulmus alata, showing corky bark-ridges
• 
  Leaves of Ulmus alata

Like the other North American species of elm, U. alata is very susceptible to Dutch elm disease and Elm Yellows (Elm phloem necrosis).^[8]

Ulmus alata is rarely cultivated beyond its natural range. It remains in commercial production in the US, and is occasionally available in Europe. At the beginning of the 19th century, the tree was one of the three American elm species cultivated in ornamental plantations in Britain,^[9] but is now rare there. Several specimens are grown in New Zealand.^[10]

On the silty uplands of the Mississippi Delta, Ulmus alata can attain 27 m (89 ft) in height, although the trunk diameter rarely exceeds 60 cm (24 in) d.b.h. In the old growth Fernbank Forest in Atlanta, Georgia, the species attains heights up to 39 metres (128 ft). A tree measuring 40 metres (130 ft) high has been reported from the Congaree National Park in South Carolina.^[11] However, the USA National Champion, measuring 27 metres (89 ft) high in 2009, grows in Hopewell, Virginia.^[12][3]

• Lace Parasol

Ulmus alata is of minimal commercial significance, its hard timber considered no more remarkable than that of other American elms, and of limited use because of the commonly small size of the trees. However, owing to its resistance to splitting, it is used to make high-quality hockey sticks.^[13]

North America

• Arnold Arboretum, US. Acc. no. 404-95, wild collected.
• Bartlett Tree Experts,^[14] US. Acc. no. 1438, unrecorded provenance.
• Brooklyn Botanic Garden,^[15] New York City, US. Acc. nos. 730275, X00886
• Bernheim Arboretum and Research Forest Bernheim Arboretum and Research Forest - Connecting People with Nature, Clermont, Kentucky, US. No details available.
• Morton Arboretum, US. Acc. no. 116-96, wild collected Papoose Lake, Illinois.^[16]

Europe

• Brighton & Hove City Council, UK. NCCPG Elm Collection.^[17] One tree at East Brighton Park, UK champion 13 m (43 ft) high, 31 cm (12+1⁄4 in) d.b.h. in 2001.^[18]
• Grange Farm Arboretum, Lincolnshire, UK. Acc. no. 506
• Royal Botanic Garden Edinburgh, UK. Acc. no. 20080092, from seed wild collected in USA.^[19]
• Thenford House, Northamptonshire, UK. No details available.

Australasia

• Manukau Cemetery & Crematorium, Auckland, New Zealand. No details available.

North America

Widely available.

Europe

• Arboretum Waasland Archived 19 February 2012 at the Wayback Machine, Nieuwkerken-Waas, Belgium.
• Grange Farm Plants, Spalding, Lincolnshire, UK.
• Plantentuin Esveld, Netherlands.

• Sheffield's Seeds Co. Inc., New York, US.

• Snow, G. A. (1990). "Ulmus alata". In Burns, Russell M.; Honkala, Barbara H. (eds.). Hardwoods. Silvics of North America. Vol. 2. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA) – via Southern Research Station.
• U. alata, Flora of North America, www.efloras.org