Ursinae

Ursinae; Temporal range: Barstovian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	A Eurasian brown bear (Ursus arctos arctos)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Ursidae
Subfamily:	Ursinae; Fischer de Waldheim, 1817
Genera
	See text.

Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998), and Jin et al. (2007).^[1]^[2]^[3]

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The genera Melursus and Helarctos are sometimes also included in Ursus. The Asiatic black bear and the polar bear used to be placed in their own genera, Selenarctos and Thalarctos; these are now placed at subgenus rank.

Subfamily Ursinae Fischer de Waldheim, 1817
- †Aurorarctos Jiangzuo & Flynn, 2020
  - †Aurorarctos tirawa Jiangzuo & Flynn, 2020
- Helarctos Horsfield, 1825
  - Helarctos malayanus (Raffles, 1821) – sun bear
  - †Helarctos sinomalayanus (Thenius, 1947)
- Melursus Meyer, 1793
  - Melursus ursinus (Shaw, 1791) – sloth bear
  - †Melursus theobaldi (Lydekker, 1884)
- †Protarctos Kretzoi, 1945
  - †Protarctos abstrusus (Bjork, 1970)
  - †Protarctos boeckhi (Schlosser, 1899)
  - †Protarctos ruscinensis (Depéret, 1890)
  - †Protarctos yinanensis (Li, 1993)
- Ursus Linnaeus, 1758
  - Ursus americanus (Pallas, 1780) – American black bear
  - Ursus arctos Linnaeus, 1758 – brown bear
  - †Ursus deningeri Richenau, 1904
  - †Ursus dolinensis (Garcia & Arsuaga, 2001)
  - †Ursus etruscus Cuvier, 1823
  - †Ursus ingressus Rabeder, Hofreiter, Nagel & Withalm 2004
  - †Ursus kudarensis Baryshnikov, 1985
  - Ursus maritimus Phipps, 1774 – polar bear
  - †Ursus minimus (Devèze & Bouillet, 1827)
  - †Ursus pyrenaicus (Depéret, 1892)
  - †Ursus rossicus Borissiak, 1930
  - †Ursus sackdillingensis Heller, 1955
  - †Ursus savini (Andrews, 1922)
  - †Ursus spelaeus Rosenmüller, 1794 – cave bear
  - Ursus thibetanus (Cuvier, 1823) – Asiatic black bear
  - †Ursus vitabilis? Gidley, 1913

A number of hybrids have been bred between American black, brown, and polar bears (see Ursid hybrids).

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[1]
Bjork, Philip R. (1970). "The Carnivora of the Hagerman Local Fauna (Late Pliocene) of Southwestern Idaho". Transactions of the American Philosophical Society. 60 (7). American Philosophical Society: 3–54. doi:10.2307/1006119. JSTOR 1006119.
[2]
Hunt, R. M. (1998). "Ursidae". In Jacobs, Louis; Janis, Christine M.; Scott, Kathleen L. (eds.). Evolution of Tertiary Mammals of North America: Volume 1, Terrestrial Carnivores, Ungulates, and Ungulate like Mammals. Cambridge, UK: Cambridge University Press. pp. 174–195. ISBN 0-521-35519-2.
[3]
Jin, C; Ciochon, RL; Dong, W; Hunt Jr, RM; Liu, J; Jaeger, M; Zhu, Q (2007). "The first skull of the earliest giant panda". Proceedings of the National Academy of Sciences of the United States of America. 104 (26): 10932–7. Bibcode:2007PNAS..10410932J. doi:10.1073/pnas.0704198104. PMC 1904166. PMID 17578912.

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[1] [1]
Bjork, Philip R. (1970). "The Carnivora of the Hagerman Local Fauna (Late Pliocene) of Southwestern Idaho". Transactions of the American Philosophical Society. 60 (7). American Philosophical Society: 3–54. doi:10.2307/1006119. JSTOR 1006119.

[2] [2]
Hunt, R. M. (1998). "Ursidae". In Jacobs, Louis; Janis, Christine M.; Scott, Kathleen L. (eds.). Evolution of Tertiary Mammals of North America: Volume 1, Terrestrial Carnivores, Ungulates, and Ungulate like Mammals. Cambridge, UK: Cambridge University Press. pp. 174–195. ISBN 0-521-35519-2.

[3] [3]
Jin, C; Ciochon, RL; Dong, W; Hunt Jr, RM; Liu, J; Jaeger, M; Zhu, Q (2007). "The first skull of the earliest giant panda". Proceedings of the National Academy of Sciences of the United States of America. 104 (26): 10932–7. Bibcode:2007PNAS..10410932J. doi:10.1073/pnas.0704198104. PMC 1904166. PMID 17578912.

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Ursinae

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Classification

References

Ursinae Temporal range: Barstovian–Recent PreꞒ Ꞓ O S D C P T J K Pg N

A Eurasian brown bear (Ursus arctos arctos)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Ursidae
Subfamily:	Ursinae Fischer de Waldheim, 1817
Genera
See text.