International Code of Phylogenetic Nomenclature (PhyloCode) 1st Edition Philip D. Cantino and Kevin de Queiroz All Chapter Instant Download
International Code of Phylogenetic Nomenclature (PhyloCode) 1st Edition Philip D. Cantino and Kevin de Queiroz All Chapter Instant Download
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International Code
of Phylogenetic
Nomenclature
(PhyloCode)
International Code
of Phylogenetic
Nomenclature
(PhyloCode)
Version 6*
* This is the first version published in hard copy. Previous versions are available at www.phylocode.org.
** Authors listed alphabetically
CRC Press
Taylor & Francis Group
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Boca Raton, FL 33487-2742
© 2020 by Taylor & Francis Group, LLC
CRC Press is an imprint of Taylor & Francis Group, an Informa business
No claim to original U.S. Government works
First published in digital version April 29, 2020.
International Standard Book Number-13: 978-1-138-33282-9 (Paperback)
International Standard Book Number-13: 978-1-138-33286-7 (Hardback)
This book contains information obtained from authentic and highly regarded sources. Reasonable efforts have been made
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Preface.........................................................................................vii
Preamble................................................................................. xxxix
Division I. Principles......................................................................1
Division II. Rules...........................................................................3
Chapter I. Taxa......................................................................5
Article 1. Categories of Taxa......................................5
Article 2. Clades........................................................5
Article 3. Hierarchy and Rank...................................7
Chapter II. Publication..........................................................9
Article 4. Publication Requirements...........................9
Article 5. Publication Date......................................11
Chapter III. Names.............................................................13
Article 6. Categories of Names.................................13
Article 7. General Requirements for Establishment.....15
Article 8. Registration..............................................16
Chapter IV. Establishment of Clade Names.........................21
Article 9. General Requirements and
Phylogenetic Definitions...........................21
Article 10. Selection of Clade Names for
Establishment...........................................40
Article 11. Specifiers and Qualifying Clauses............51
Chapter V. Selection of Accepted Clade Names...................67
Article 12. Precedence...............................................67
Article 13. Homonymy..............................................68
v
Contents
vi
Preface
vii
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viii
Preface
ix
Preface
taxonomies. (2) All taxa named under this code are clades. Clades
are products of evolution that have an objective existence regardless
of whether they are named. As a consequence, once a clade is named
and its name associated with a phylogenetic definition, its composi-
tion and diagnostic characters become questions to be decided by
empirical evidence rather than by personal decisions. (3) In addi-
tion to applying names to nested and mutually exclusive taxa, as in
traditional nomenclature, the phylogenetic system allows names to
be applied to partially overlapping clades. This provision is neces-
sary to accommodate situations involving clades of hybrid origin.
(4) In contrast to the rank-based codes, which use (implicit) defi-
nitions based on ranks and types to determine the application of
names, phylogenetic nomenclature uses explicit phylogenetic defini-
tions. Species, specimens, and apomorphies cited within these defi-
nitions are called specifiers because they are used to specify the clade
to which the name applies. These specifiers function analogously to
the types of rank-based nomenclature in providing reference points
that determine the application of a name; however, they differ from
types in that they may either be included in or excluded from the
taxon being named, and multiple specifiers may be used. (5) The
fundamental difference between the phylogenetic and rank-based
systems in how names are defined leads to differences in how syn-
onyms and homonyms are determined in practice. For example,
under the PhyloCode, synonyms are names whose phylogenetic defi-
nitions specify the same clade, regardless of prior associations with
particular ranks; in contrast, under the rank-based codes, synonyms
are names at the same rank whose types are included within a single
taxon at that rank, regardless of prior associations with particular
clades. (6) Another novel aspect of the PhyloCode is that it permits
taxonomists to restrict the application of names with respect to clade
composition. If a taxonomist wishes to ensure that a name refers to
a clade that either includes or excludes particular taxa, this result
may be achieved through the use of additional internal or external
specifiers (beyond the minimal number needed to specify a clade), or
x
Preface
xi
Preface
clades much more easily than they can under the rank-based codes.
For many researchers, naming clades is just as important as nam-
ing species. In this respect, the PhyloCode reflects a philosophical
shift from naming species and subsequently classifying them (i.e.,
into higher taxa) to naming both species and clades. This does not
mean, however, that all clades must be named. The decision to name
a clade (or to link an existing name to it by publishing a phyloge-
netic definition) may be based on diverse criteria, including (but not
restricted to), level of support, phenotypic distinctiveness, economic
importance, and whether the clade has historically been named.
Another benefit of phylogenetic nomenclature is that it permits
(though it does not require) the abandonment of categorical ranks,
which would eliminate the most subjective aspect of traditional tax-
onomy. The arbitrary nature of ranking, though acknowledged by
most taxonomists, is not widely appreciated by non-taxonomists.
The existence of ranks encourages researchers to use taxonomies
inappropriately, treating taxa at the same rank as though they were
comparable in some biologically meaningful way—for example,
when they count genera or families to study past and present pat-
terns of biological diversity. A rankless system of taxonomy, which is
permitted but not required by the PhyloCode, encourages the devel-
opment of more appropriate uses of taxonomies in such studies, such
as counting clades or species that possess properties relevant to the
question of interest, or investigating the evolution of those proper-
ties on a phylogenetic tree.
An advantage of the PhyloCode over the rank-based codes is that
it applies at all levels of the taxonomic hierarchy. In contrast, the
zoological code does not extend its rank-based method of definition
above the level of superfamily, and the botanical code extends that
method of definition only to some names above the rank of family
(automatically typified names) and the principle of priority is not
mandatory for those names. Consequently, at higher levels in the
hierarchy, the rank-based codes permit multiple names for the same
taxon as well as alternative applications of the same name. Thus, as
xii
Preface
xiii
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xiv
Preface
xv
Preface
xvi
Preface
xvii
Preface
xviii
Preface
xix
Preface
xx
Preface
xxi
Preface
xxii
Preface
xxiii
Preface
xxiv
Preface
xxv
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armies were in the field, and the quick and signal triumph of
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the American people a new and noble responsibility.
{651}
"We favor home rule for and the early admission to Statehood
of the Territories of New Mexico, Arizona and Oklahoma.
{652}
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declarations in its platform as to any questions concerning
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{653}
"We declare that there are but two real parties to-day
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endurance for this Republic."
{655}