Rmrs gtr077
Rmrs gtr077
Rmrs gtr077
Department
of Agriculture High Mountain Lake
Forest Service
Rocky Mountain
Research Natural
Research Station
General Technical
Areas in Idaho
Report
RMRS-GTR-77-CD
Fred W. Rabe
June 2001
High Mountain Lake
Research Natural Areas
in Idaho
Fred W. Rabe
The Author
Abstract
Fred W. Rabe
Contents
Introduction..............................................................................................................................................1
Classification ...........................................................................................................................................9
Sawtooths Subregion..........................................................................................................................123
Surprise Valley Lake and Pond.................................................................................................125
Smiley Mountain Lake and Ponds............................................................................................129
Broad Valleys Subregion.....................................................................................................................137
Kenney Creek Pond .................................................................................................................139
Upper Mill Lake.........................................................................................................................141
Upper Merriam Lake.................................................................................................................145
Recommendations...............................................................................................................................166
Table 2. Association between physical, chemical and biotic factors with production potential in
high lakes ..................................................................................................................................10
Table 4. Classification elements associated with high lakes and ponds ..............................................162
Table 5. Dominant Ephemeroptera, Plecoptera and Trichoptera from lakes ponds and
streams....................................................................................................................................166
Glossary ...............................................................................................................................................169
1
G
F
E B
D C
A
A. alpine cirque lake
B. subalpine ponds
C. wet meadow
D. meandering stream
E. whitebark pine
F. mixed subalpine fir -Englemann spruce forest
G. heather meadows
From Rabe and Savage (1977).
Research Natural Areas consist of a number of cells or basic ecological units. Mill Lake RNA in the Salmon
-Challis National Forest comprises an aggregation of cells. Natural area needs assessment identifies cells
that are under-represented in the system (Chadde et al. 1996 and Rust 2000).
2
Purposes of Natural Areas
3
Over a period of years in the high Uinta Mountains of Utah The baseline data from Upper Steep Lake provided an impor-
Rabe completed age and growth studies of brook trout, tant control enabling the investigators to see that a slight
Salvelinus fontinalis (Rabe 1968), a transplantation study of increase in nitrates and a decrease in zooplankton occurred
brook trout (Rabe 1967b) and an age and growth study of after the eruption. These effects were only temporary. (Rabe
rainbow trout,Oncorhynchus gairdneri (1967a). In addition 1982).
an artificial fertilization experiment was performed in sever-
al of the waters (Rabe 1969). These lakes in the Swift Creek In the 1970s, some high lakes in Idaho and Montana were
drainage of the Uintas are protected as wilderness. used as baseline sites to monitor the effects of acid rain.
Since a majority of these pristine waters are nutrient poor,
Reference Areas have relatively low pH values and a poor buffering capacity,
Natural areas may provide baseline or reference areas. any effects of acid rain would be easily detected compared
These sites enable us to better understand our impact on to lower lakes and streams with harder waters and more
the world and it upon us by studying natural systems and impacted conditions.
then comparing them to systems that have been affected by
human activities. Classrooms
Natural areas may serve as teaching sites or outdoor labo-
Natural areas provide reference points that we can use to ratories where we can learn more about the natural world.
measure the structure and function of ecosystems. They They are as important to field scientists as a physiology lab
help define ranges of natural variability and present a clear- might be to a medical student; hence these valuable
er understanding of natural processes at work in the larger resources should be protected. We might think of natural
landscape. In addition, they provide an early warning sys-
areas as a library with each individual natural area a book,
tem for environmental problems. The same natural laws that
so both students and scientists can read the lives of
govern fish or plankton in a lake govern us.
streams, lakes, wetlands, trees, flowers, soils and land-
The Sawtooth National Forest has been targeting high lakes forms.
in the Sawtooth Wilderness as monitoring sites (Moseley
1996). Fish, amphibians and plants are sampled in five year Belvidere Lakes RNA on the Payette National Forest was
cycles to determine long-term trends of the baseline data the classroom setting for one group of students participating
measured. A floristic checklist combined with a measure of in a University of Idaho ecology class in 1978. Students
relative abundance of each species in the lakes and asso- hiked several miles up a steep grade carrying a microscope,
ciated wetlands is the goal of this project. folding table, glassware, rubber rafts and sundry other sam-
pling equipment. The objective was to acquaint students
with the methods of sampling high mountain lakes and ana-
lyzing the data.
Biotic Preserves
Natural areas may act as preserves for biotic diversity and
A trip to Lower Steep Lake (Clearwater National Forest) processes. The sites contain both representative and rare
was made to measure any impact Mount St. Helen ash species of plants and animals. It is difficult to imagine study-
fall (gray in color) might have had on plankton. Most of
ing nature without these pristine habitats in which to work.
the lake was frozen, with ash covering the surface.
As with libraries, our understanding of ecological systems in
Steve Crumb compared invertebrate communities of Upper natural areas is enhanced when we have a catalog of native
and Lower Steep Lakes in the Bitterroot Range (Crumb flora and fauna and a description of the habitat.
1977). Mount St. Helens erupted soon after he completed
his study. Since chemical and biological data for the lakes The following examples demonstrate how studies in RNAs
existed before the eruption, the Forest Service asked Rabe have revealed unique or unusual characteristics of high
to determine what effect, if any, the ash fall might have had lakes. While studying plant composition in glacial lakes in
on the water chemistry and biota in one of the lakes. the Sawtooths, Moseley (1996) observed a sedge and a
rush that while common in low elevation peatlands are
rarely observed at higher elevations.
4
Evenden, A. G.; Moeur, M.; Shelly, S.; Kimball, S. F.;
Wellner C. A. ( In Press). Forest Service Research Natural
Areas in the Northern Rocky Mountains and Intermountain
West.U.S. Department of Agriculture, Forest Service,
RMRS.
Lower Steep Lake (North Fork of the Clearwater drainage) Parr, W. H.; Rabe, F. W. 1973. A cutthroat population in a
is the only lake north of the Salmon River that contains a small subalpine meadow lake. Idaho Academy of Science 9:
breeding population of golden trout (Salmo aquabonita). 44-48.
The only known case of a freshwater shrimp (Gammarus Parr, W. H.; Rabe, F. W.; Wissmar, R. C.1968. Investigations
lacustris) in high lakes north of the Salmon River was col- of subalpine lakes, Five Lakes Butte, Idaho. Journal of
lected in Upper Steep Lake which contains no fish. Fishless Idaho Academy of Science 8:1-5.
lakes and ponds are important because they provide a habi-
tat for certain invertebrates, amphibians and salamanders Rabe, F. W. 1967a. Age and growth of rainbow trout in four
that might not be there if fish were present. alpine lakes. Northwest Science 41 (1): 12-22.
The Forest Service in collaboration with other agencies has Rabe, F. W. 1967b. The transplantation of brook trout in an
established a national network of natural areas that repre- alpine lake. Progressive Fish Culturist 29 (1): 53-55.
sents terrestial, wetland, stream and lake ecosystems
across the country. These sites provide outdoor laboratories Rabe, F. W. 1968. Brook trout populations in high lakes.
for research and education and serve as sites for long term Northwest Science 42 (1): 20-28.
monitoring. It is now possible for scientists to cross-refer-
ence information in RNAs, Wilderness Areas, National Rabe, F. W. 1969. Artificial fertilization of a small high alti-
Parks, private nature preserves, botanical areas and State tude lake. American Midland Naturalist 81 (1): 281-284.
heritage areas. This library of knowledge will grow as we
acquire new sites and study them more thoroughly. Rabe, F. W.; Savage, N. L. 1977. Aquatic natural areas in
Idaho. Moscow: Idaho Water Resources Research Institute.
Literature Cited University of Idaho. 109 p.
Bahls, P. ; Stickney, M. B. 1987. Preliminary report of the high Rabe, F. W. 1982. Effects of volcanic ash on plankton and
lake fisheries project on the Moose Creek Ranger District. Nez benthos in subalpine lakes. Unpublished paper on file at
Perce National Forest. Grangeville, ID. 234 p. U.S. Department of Agriculture, Forest Service, Rocky
Mountain Research Station, Ogden, UT.
Bahls, P. 1990. Ecological implications of trout introductions
to lakes of the Selway Bitterroot Wilderness, ID. Corvallis, Rabe, F. W.; Breckenridge, R. M. 1985. Physical and chem-
OR: Oregon State University. 55 p. Thesis. ical factors of glacial lakes in Northern Idaho as related to
area selection. Journal of Idaho Academy of Science 21 (1-
Bahls, P. 1992. The status of fish populations and manage- 2): 1-15.
ment of high mountain lakes in the Western United States.
Northwest Science 66 (3): 183-193. Rust, S. K. 2000. Representativeness assessment of
research natural areas on National Forest System lands in
Crumb, S. 1977. Long term effects of fish stocking on the Idaho. Gen. Tech. Rep. GTR-45. Fort Collins, CO: U.S.
invertebrate communities of Steep Lake, ID. Moscow, ID: Department of Agriculture, Forest Service, Rocky Mountain
University of Idaho. 27 p. Thesis. Research Station. 129 p.
Espinosa, F. A.; Orme, M. L.; Dolan, J. J. 1977. A limnolog- Thorson, D. M.1979.Sandpoint mountain lake management
ical survey of subalpine lakes in the Buffalo Hump Area. plan and inventory. U.S. Department of Agriculture, Forest
U.S. Department of Agriculture, Forest Service, Nez Perce Service, Sandpoint Ranger District, Panhandle National
National Forest, Grangeville, Idaho. 183 p. Forests, Coeur d’Alene, ID.
5
Wissmar, R. C.; Rabe, F. W. 1970. Crustacean populations
and sampling techniques in four mountain lakes of Idaho.
Transactions American Microscopical Society 89 (2): 205-
215.
6
FIELD METHODS
From a high point on shore an attempt was made to observe
High mountain lake RNAs are typically located in remote dominant bottom substrate in the littoral zone, that area in
areas. Access to these sites is almost always via rugged hik- the lake less than 3 m in depth. Substrates were recorded
ing trails making field work especially challenging. In addition as silt, sand, gravel, rubble, boulders, bedrock or a combi-
to packing in supplies and camping equipment, the nation of the above.
researchers carried sampling gear. The sampling equipment
was lightweight, durable and compact weighing less than 15 Lake sampling procedure
pounds. Distance and time constraints sometimes limited the
number of sampling activities. Field collections were made Collecting samples from the lake posed a challenge.
from the late 1960s through 1999. Investigators needed a boat or flotation device that was sta-
ble and sturdy enough to allow them to safely navigate the
Shoreline sampling procedure lakes; at the same time, the craft had to be light enough to
carry to the research site. The Curtis Designs ultralight
Some field methods were modified from Bahls (1989). inflatable boat weighed only 20 ounces (see below). It is
Sections of shoreline too steep to hike safely were observed made of urethane-coated nylon taffeta with heat sealed
from high points above the lake and from the inflatable boat. seams. A Therm-a-Rest air mattress served as a seat and
The percent coverage of terrestrial vegetation types two ultralight paddles similar to ping-pong paddles were
(trees, shrubs, herbs) in a 10 m wide strip around the lake used to move the raft.
were noted together with the percentage of open ground
coverage (talus and rock).
(depending upon the depth of the lake) into the water and
retrieved rapidly to sample a vertical haul of zooplankton.
7
An estimate of the dominant bottom substrate and per- Fish were collected from the raft by rod and reel.
cent littoral zone was attempted from the boat. In addition The data used in this report were collected by different
the researcher estimated percent bottom coverage of any researchers over several years. Ideally, the data record
submergent macrophytes. would contain information on morphometry, water chem-
istry, geology, macrophytes, zooplankton, macroinverte-
A bathymetric survey of the lake was attempted in lakes less brates and vertebrates for every lake. Not all of this infor-
than 4 hectares (10 acres). The study lake is photo-copied mation is recorded for every lake but the databases can be
from a 7.5 minute USGS topographic map onto a transparen- updated as continued research adds to our body of knowl-
cy. An overhead projector was used to enlarge the image of edge. RNAs targeted as natural “classrooms” can have the
the lake for tracing onto water resistant paper (Bahls 1989). dual benefit of providing hands-on research experience for
students and providing valuable data for the high mountain
lakes RNA library.
8
CLASSIFICATION rocky basin but are not above tree line. Montane lakes are
found lower at mid-mountain elevations.
It is important to choose sites that provide a wide range of
Size-depth
ecosystem diversity as RNAs. The classification scheme
developed for high mountain lake RNAs is based on previ-
ous research by the author and others as discussed in the Classifying water bodies according to size can be difficult. In
introduction. The system takes into account factors such as this report they are identified as lakes or ponds. Since the
lake form, chemistry, and biotic characteristics ensuring that terms “pond” and “lake” defy description (Reid 1961), the sites
RNAs represent a diverse range of ecosystems. were categorized according to the above graph which takes
into consideration depth and area (Anderson 1971).
According to Wuerthner (1986), there are 81 mountain
ranges in the state; only the Snake River Plain and some The term “lake” has some additional sub-categories. Small
large prairies and valleys lack hills and mountains. The high lakes are defined as those less than 4 hectares (10 surface
lake RNAs cover seven mountainous subregions deter- acres). Large lakes exceed 4 hectares in size (Bahls
mined by similarities of geology, terrain, climate and plant 1989). Deep lakes are over 4 meters (15 ft) in depth, and
cover (Wuerthner 1986). shallow lakes are less than 4 meters.
Elevation
Production Potential
9
Table 1. Classification elements of high mountain lakes
Table 2. Association between physical, chemical, and biotic factors with production potential in high lakes
Dominant bottom substrate bedrock, boulders soft sediments cobble, rubble, gravel
in littoral zone
10
Johnston found that where these parameters were relative-
ly low, fish required a longer time to reach a certain length
than when these values were higher. Since high lake condi-
tions are believed to be somewhat similar in Idaho and
Washington, Johnston’s study was applied to selected mor-
phometric, chemical and biotic conditions in Idaho lakes
(Rabe and Breckenridge 1985).
The non-weighted values for all of the factors were then added
together to get an overall score, and the lake was assigned a
high, medium-high, medium, medium-low or low production
potential based on that score. The numeric ranges were 11-14
for high production potential, 9-10 medium-high, 7-8 medium,
6 medium-poor and 1-5 for poor production potential.
The percentage littoral zone in
Aspect this steep-sided lake is small.
11
Alkalinity
Shoreline development
Sedge beds
12
.
(pH 5.5-7.4)
(pH 5.5-7.4)
(pH 7.5-8.4)
13
within the five classification categories. The modifiers are Geomorphic lake types in this study are patterned after
pH, rock type and hydrology (Table 3). Lake water pH is Rabe and Breckenridge (1985).This classification describes
measured in the field. Rock types for some RNAs were the lake basins using very general terminology and com-
described by the Conservation Data Center Site Basic pares them using common parameters. Six types of lakes
Records and personnel from Idaho Geological Survey. are recognized (Table 1).
Sedge bed
Cirque lake
Cirque -scour
Cirque-scour lakes form when alpine glaciers scour out a
basin down-valley from the headwall. These lakes usually
occupy basins carved in less resistant bedrock caused by a
variation in geologic structure or rock composition. The
basins are often the result of multiple glaciations and repre-
sent different ice advances.
Lake Origin
Paternoster lakes are a variety of cirque-scour lakes that Upland lakes form in depressions scoured by ice caps on
form a linear chain or string of “bead lakes” connected by a gently rolling or upland valleys. This type might represent
stream in a glacial valley. These lakes are usually separated early Wisconsin ice cap glaciation or an even older pre-
by steep drops or by moraines from different ice advances. Wisconsin period.
Upland lake
Beaver modified
Paternoster lakes
Moraine
15
Such lowland lakes are often classed as semidrainage Savage, N. L.; Rabe, F. W. 1979. Stream types in Idaho: An
(Pennak 1969). Semidrainage lakes lack a permanent out- approach to classification of streams in natural areas
let. They are small in size, shallow, have a high organic con- Biological Conservation 15: 301-315.
tent and are usually located in a meadow (meadow lake).
Semidrainage lakes in this study were montane or subalpine Wellner, C. A.; Moseley, C. A. 1986. Establishment record
in elevation and were mostly cirque-scour in origin. for Theriault Lake Research Natural Area within Panhandle
National Forest, Shoshone County, ID. U. S. Department of
Literature Cited Agriculture, Forest Service. On file at Intermountain Region,
Ogden, UT. 18 p.
Anderson, R. S. 1971. Crustacean plankton of 146 alpine
and subalpine lakes and ponds in Western Canada. Journal Wuerthner, G. 1986. Idaho Mountain Ranges. Idaho
Fisheries Research Board of Canada 28: 311-321. Geographic Series. Helena, MT: American Geographic
Publishing. 101 p.
Bahls, P. 1989. Preliminary report of survey methodology for
high mountain lakes. High Lake Fisheries Project. U.S.
Department of Agriculture, Forest Service. Nez Perce
National Forest, Grangeville, ID and Idaho Fish and Game
Department, Lewiston, Idaho. 50 p.
18
Description of high mountain lake Research The individual descriptions of RNA lakes and ponds are
arranged in the following format.
Natural Areas in Idaho
Introduction: The introduction contains information about
The ten subregions in the state are determined by similari- the number of water bodies in each RNA, which ones were
ties of geology, terrain, climate and plant cover. RNA high sampled and by whom. The sampling date is also included.
lakes described in this book are located in seven of these
regions. A map of each of the subregions (see example Location: This section contains the description of the geo-
below) preceeds the description of RNAs. graphic area, the ecoregion section, county, and the name
of the USGS quad map. Although we have chosen to use
the National Hierarchical Framework of Ecological Units
(McNab and Avers 1994; Bailey 1995) in the publication, it
is important to note that refinements in this classification
have been made and are being applied within certain geo-
graphic areas. For example, the State of Idaho is utilizing a
revised version of the national hierarchy that was developed
for the Interior Columbia Basin Ecosystem Management
Project (Menakis and Long 1996; Rust 2000).
Field collections were made from the 1960s through 1999. Zooplankton: Identification of copepods and cladocera are
Distance and time constraints sometimes limited the num- listed.
ber of sampling activities. In addition the data used in this
report were collected by different researchers over the Macroinvertebrates: Macroinvertebrates, predominantly
years. Even though information about some of these lakes aquatic insects, are identified from each water body.
is not currently available, the databases can be updated as Crustaceans, snails, clams, aquatic earthworms and leech-
continued research adds to our body of knowlege. es are also listed. Macroinvertebrates are described from a
lake inlet or outlet.
17
Vertebrates: When observed, fish, amphibians and sala-
manders are described.
18
19
20
SNOWY TOP LAKE Geology
21
Classification
19.5
18
12
The lake depth drops off abruptly from shore thus the littoral zone
is limited. The soft bottom is covered with a layer of felt-like algae.
Logs have accumulated near the outlet end of the lake. The out-
let stream late in September was partially dried out making it dif-
ficult for fish to gain access from the lake. The stream substrate Snowy Top Lake and view of Snowy Top Mountain summit. The
consists of various sized rock coated with algae. shoreline of the lake is ringed with conifers and alder trees.
22
Logs piled up at outlet side of Snowy Top Lake.
Vegetation
23
Vertebrates
Paraleptophlebia debilis
Sketch credit: McCafferty 1983
Outlet stream
Ephemeroptera
Epeorus deceptivus
Paraleptophlebia debilis
Trichoptera
Polycentropus sp.
Chironomidae
24
Three Ponds western edge of the Purcell Trench which was scoured out
by continental ice sheets. Moseley and Wellner (1987)
report the ponds were formed from glacial scouring that
Three Ponds Research Natural Area exposed the bare granitic ridges. This is especially evident
Idaho Panhandle National Forest north of the ponds. A deep valley marks a strike-slip fault on
the west side of the RNA..
Three ponds are located in the RNA. Charles Wellner and
Fred Rabe hiked to the site on June 26, 1982. Aquatic sam-
pling occurred at the West Pond.
Location
West Pond
• Montane, small, shallow, upland pond
• Medium-high production potential
• Circumneutral water in a granite basin
East • Inlet: ephemeral; Outlet: ephemeral
Middle Pond
Pond
Vegetation
West Pond
Aeshnidae Libelluidae
Porifera Sphaeridae
Physidae Caenidae
Phryganeidae Limnephilidae
Glossiphonidae Phyryganeidae
Dytiscidae Gerridae
Chaoboridae Coenagrionidae
Chironomidae Planorbidae
Western paper birch (Betula An unidentified frog species was observed. No fish were
papyrifera ) noted. A more thorough collection of macroinvertebrates
and amphibians from the ponds is recommended.
Literature Cited
26
Scotchman No. 2 Pond
Pond
27
Literature Cited
28
POND PEAK POND
Pond Peak Research Natural Area
Idaho Panhandle National Forest
Location
Wallace Formation of the Belt Supergroup. From: Wellner
The RNA is located in the Shoshone Range in the Idaho and Moseley (1988).
Panhandle northeast of Interstate 90 near Kingston, Idaho.
The pond drains into Shoshone Creek and from there into
the North Fork of the Coeur d’Alene River.
Classification
Geology
29
Aquatic physical-chemical factors Vegetation
Lake surface area (hectares): 0.4 (0.9 acres) Old growth stands consisting almost exclusively of moun-
Length of shoreline (m): 231 (758 ft) tain hemlock (Tsuga mertensiana) in the climax stage of
Maximum depth (m): 1.2 (4 ft) succession are one of the main reasons this area was
Elevation (m): 1649 (5410 ft) selected as an RNA.
Aspect: NE
Percent shallow littoral zone: 100 Sitka alder (Alnus sinuata), sedges (Carex spp.) and
Dominant bottom substrate: soft sediments conifers occur along 50 percent of the shoreline. The alder
Shoreline development: 1.06 and sedges are concentrated on the east shore (Wellner
Lake edge %: talus -50, conifers-20, shrubs-15, sedges-15 and Moseley 1988).
Alkalinity (mg/l): 4
Conductivity (micromhos/cm): 3
pH: 6.6
Inlet: seep
Outlet: none
Zooplankton
30
Polyphemus pediculatus is a common zoo-
plankton in shallow weedy water. It is the
sole genus in the family Polyphemidae.
Sketch from Brooks (1963).
31
Wellner, C. A. and R. K. Moseley 1988. Establishment record
for Pond Peak Research Natural Area, Shoshone County. U.S.
Department of Agriculture, Forest Service. Unpublished report
on file at Northern Region, Missoula, MT.19 p.
32
33
34
Theriault Pond
Location
From Clarkia take FS Road 301 east for one mile. Then
travel FS Road 321 north over Hobo Pass down to Marble
Creek. Continue on 321 until it joins with FS Road 216, 16 Classification
miles from Clarkia. Take FS Road 216 east for about 9 miles
bearing left at junctions until reaching FS Road 1936. • Subalpine, small, shallow cirque pond
Proceed northwest on FS 1936 for 4.5 miles until road • Medium-high production potential
junctions with FS Road 1938. Continue on FS Road 1938 • Circumneutral water in Precambrian Belt rock
until you come to the end of the road. Hike up the trail sev- • Inlet: stream; Outlet: Meandering glide stream
eral hundred yards to the top of Marble Mountain. Proceed
down steep hillside to Theriault Pond.
Geology
36
Vegetation
Polyphemus pediculus
Suborder Harpacticoida
Chydorus sp.
37
Macroinvertebrates
A Coeur d’Alene salamander (Plethodon idahoensis) was
observed about 25 m from the lake shore in a wet area. The
Pond edge Outlet range of this vertebrate is somewhat restricted; it is one of
Trichoptera the less well known amphibians in our region (Corkran and
Homophylax sp. X Thoms 1996).
Protoptila sp. X
Ephemeroptera
Siphlonurus sp. X
Odonata
Libelluidae X
Family Cordulegastridae X
Ischnura / Enallagma X
Coleoptera
Ilybius / Agabus X X
Hemiptera Spotted frog (Rana pretiosa) was extremely com-
Gerris sp. X mon in both pond and streams. From Corkran and
Thoms (1996)
Notonecta kirbiyi X
Diptera
Subfamily Tanypodinae X X
Family Psychodidae X
Family Culicidae X
Pelecypoda
Pisidium sp. X X
Literature Cited
Fred Rabe, Bob Wissmar and Bill Parr studied ten lakes
including Lower Bacon Lake in the Five Lakes Butte area in
1968. Upper Bacon Lake was not sampled. Upper and
Lower Bacon Lakes were established within Five Lakes
Butte RNA in 1986 after a visit to the site by Charles
Wellner and Bob Moseley. The following report deals with
research on Lower Bacon Lake and several other lakes in
the Five Lakes Butte area.
Location
Lower
Lake
Upper
Lake
The dashed line is the original valley floor. The solid line is
USGS Quad: BACON LAKE. the valley floor after glaciation. B = basin; S = step. Parr
et al. (1968).
39
Classification
Vegetation
40
Zooplankton
Diaptomus sp.
Daphnia sp.
Holopedium gibberum
Conochilus sp.
Family Ceratopogonidae
Family Limnephlidae
Family Dytiscidae
Family Sialidae
Family Libelluidae
Family Sphaeriidae
Class Oligochaeta
41
Research
Literature Cited
Steep Lakes Research Natural Area Upper and Lower Steep Lakes are subalpine cirque lakes sit
ated on the west side of the Bitterroot Range in the headwate
Clearwater National Forest of the North Fork of the Clearwater River. The site is
Precambrian Belt quartzites and argillites that have been su
During the summer of 1997, Fred Rabe and Steve Crumb jected to alpine glaciation. Carbonaceous argillites are present
conducted research at the two Steep Lakes and pond. the basins accounting for the relatively high calcium carbona
content of the water (Crumb 1977).
Location
Lower
Pond Lake
Upper
Lake
43
Aquatic physical-chemical factors Upper Steep Lake
Lower Steep Lake Area (hectares): 2.3 (5.7 acres)
Area (hectares): 2.3 (5.8 acres) Length of shoreline (m): 625 (2051 ft)
Length of shoreline (m): 610 (2001 ft) Maximum depth (m): 4.6 (15.1 ft)
Aspect: NW Elevation (m): 2012 (6600 ft)
Maximum depth (m): 4.7 (15 ft) Aspect: N
Elevation (m): 1800 (5904 ft) Percent shallow littoral zone: 20
Percent shallow littoral zone: 20 Dominant substrate: silt and angular boulders
Dominant substrate: primarily silt and large boulders Shoreline development: 1.172
Shoreline development: 1.142 Alkalinity (mg/l): 50
Alkalinity (mg/l): 50
Inlets: 1 stream, 2 seeps
Outlet: 1 stream
The lake substrate consists of fine silt and large angular boul-
ders. Considerable log debris extends along the northwest
side and provides good cover for trout. Underwater springs
provide oxygenated water that help the trout survive in the
lake during the winter months when oxygen might be limited.
44
Upper Steep Lake is southwest of the lower lake and about
213 m (700 ft) higher in elevation. Bottom substrate consists
of silt and large boulders. Numerous springs are present.
No macrophytes and only scattered log debris occur there.
The inlet stream is approximately 3 m (10 ft) wide and lacks
the spawning gravel present in the lower lake basin. The
outlet, about 3.7m (12 ft) wide, flows into a pond that drains
down the mountainside over a steep cascade going under-
gound before reaching the lower lake.
Upper Steep Lake outlet flows into a pond. The outlet of the
pond then cascades down the mountain and goes under-
ground until entering the lower lake.
45
Vegetation
Upper and Lower Steep Lakes continued
The surrounding forests are dominated by mountain hem-
Plecoptera
lock (Tsuga mertensiana) and subalpine fir (Abies lasio-
Acroneuria sp. - stream
carpa). Whitebark pine (Pinus albicaulis) and Sitka alder
(Alnus sinuata) communities are also present (Habeck Alloperla sp. - stream
1988).
Ephemeroptera
Steep Lakes Basin was burned in 1910 and therefore has Ameletus sp.
sparse tree cover. The South Basin with older stands did not Siphlonurus sp. - stream
burn in 1910 (Habeck 1988). Little or no macrophytic vege- Cinygmula sp.
tation was noted in the lakes.
Trichoptera
Zooplankton Rhyacophila sp.
Hesperophylax sp.
Onocosmoecus sp. - stream
Diaptomus shoshone, a zooplankton, was present only in the
Psychoglypha sp.
upper lake which lacked fish (Crumb 1977). Diaptomus arapa-
hoensis was more abundant in the lower lake stocked with fish.
Megaloptera
Daphnia schodleri was more abundant in the stocked lake
Sialis sp.
except in late summer and fall when fish apparently switched
to feeding on it. Bosmina longirostris and Macrocyclops albidus
Coleoptera
were zooplankton collected in both lakes.
Gyrinus sp.
Hydrovatus sp.
Diptera
Chironomus sp.
Microspectra sp.
Procladius sp
Amphipoda
Gammarus lacustris
Rhynchobdellida
Glossiphonia complanata
Haplotaxida
Ilyodrilus templetoni
Heterodonta Freshwater shrimp (Gammarus lacustris). From
Pisidium subtruncatum Huggens et al. (1985).
.
46
Research
47
Literature Cited
48
Grave Peak Lake and Ponds
Location
Turn south onto the Elk Summit Road (FS road 360) from
about two miles east of Route 12 near the Powell Ranger
Grave Peak Pond 1 and lake viewed from the ridgeline.
Station. Follow FS road 360 about 17 miles to the junction
with FS road 358. Turn right on FS 358 and continue for two
miles to where the road has been blocked. The road Geology
becomes FS trail 45 to Kooskooskia Meadows (Wellner and
Bernatas (1991). Continue on the trail past Swamp Lake to
Grave Peak RNA is located within the Idaho batholith of
Friday Pass about 3.5 miles from the trailhead. About 100
granitic and related rocks which were uplifted and exposed
meters before reaching the top of Friday Pass, take a faint
in the late Tertiary. The area experienced multiple glacial
trail which branches to the right. It eventually takes you
events during the Quaternary, described in more detail in the
along a ridgeline separating Windy Lakes on one side and
RNA Establishment Report (Wellner and Bernatas 1991).
Grave Peak Lakes on the other. Two of the Grave Peak
water bodies can be viewed off to the right. Use extreme
caution working your way down the slope to Pond 1.
Classification
49
Aquatic physical - chemical factors
The inlet to Pond 1 flows from the talus rock for about 30
meters and then takes on a meandering glide pattern for
about the same distance to the pond. Sedges provide ripar-
ian cover. The lower section of the inlet substrate was
organic sediment. The temperature of the inlet stream is 7
degrees C.
50
Northeast view of Grave Peak Pond 2.
51
Raynold’s sedge
(Carex raynoldsii )
Hurd et al. 1998.
Macroinvertebrates
Trichoptera
Psychoglypha sp.
Allomyia sp.
Diptera
Polypedilum sp.
Rheotanytarsus sp.
After inlet stream cascades down a steep slope it levels out
for a short distance and enters Pond 2.
Grave Peak lake
Trichoptera
Clistorina sp.
Desmona sp.
Vegetation
Hesperophylax sp.
Psychoglypha sp.
Subalpine coniferous vegetation in the Graves Peak area
Megaloptera
consists of subalpine fir (Abies lasiocarpa), whitebark pine
Sialis sp.
(Pinus albicaulis), subalpine larch (Larix lyallii), and
Diptera
Engelmann spruce (Picea engelmannii).
Polypedilum sp.
Prosimulium sp.
Sedge meadows occur adjacent to the water bodies. Wellner
Rheotanytarsus sp.
and Bernatas (1991) reported yellow bog sedge (Carex dioica),
Platyhelminthes
Raynold’s sedge (Carex raynoldsii), Ross sedge (Carex rossii),
Polycelis coronata
and Drummond’s rush (Juncus drummondii).
Pelecypoda
Pisidium sp.
52
Grave Peak lake inlet
Trichoptera
Desmona sp.
Psychoglypha sp.
Diptera
Hydrobaenus sp.
Prosimulium sp
Pond 4
Grave Peak Pond 2 inlet
Trichoptera
Neothremma sp.
Plecoptera
Setvena bradleyi Pond 3
Sweltza sp.
Ephemeroptera
Nixe sp.
Ameletus sp.
Baetis bicaudatus
Diptera
Cryptolabis sp.
Subfamily Orthocladiinae Ponds 3 and 4 further downstream were not studied. Photo cred-
Simulium sp. it: Charles Wellner.
Coleoptera
Uvarus sp.
Platyhelminthes
Polycelis coronata
Lake
Pond 1
53
Literature Cited
54
Fenn Mountain Lakes The rock is Cretaceous in age and was emplaced as part of
the Idaho batholith. It is uniform in this region and the main
varying feature is the structure of the rock which can be uni-
Fenn Mountain Proposed Research Natural Area form or foliated as here, depending on the stage of metamor-
Clearwater National Forest phism (Greenwood and Morrison 1973).
Geology
55
Aquatic physical-chemical factors Literature Cited
Area (hectares 12 (29.9 acres) Bahls, P. F.1992. Report of the High Lake Fisheries Project
Length of shoreline (m): 1993 (6539 ft) Cooperative Project of Idaho Department of Fish and
Maximum depth (m): 7.5 (25 feet) Game, Region 2, Clearwater National Forest, Orofino,
Aspect: NE Idaho. 67-72.
Elevation (m): 1917 (6288 feet)
Percent shallow littoral zone: < 3 Greenwood, W. R.; Morrison, D. A. 1973. Reconnaissance
Dominant substrate: silt and rubble geology of the Selway-Bitterroot Wilderness Area. Moscow ,
Conductivity (micromhos/cm): 5 ID: University of Idaho, Bureau of Mines and Geology. l54 p.
pH: 6.6
Inlet: 13 major and 14 minor streams, 9 seeps Wellner, C. A. 1991. Letter to R. K. Moseley dated 9
Outlet: 1 stream September regarding elements occurring in Grave Peak
proposed RNA, Fenn Mountain proposed RNA and Elk
Creek proposed RNA. 2 p.
56
Salmon Mountain Lake and Ponds
Three ponds and a lake are located in the RNA. Pond 1 and
the lake were sampled by Fred Rabe on September 12-13,
1998.
Location
FS Road 468 (Magruder Corridor).
The RNAis located on the divide between the upper Selway
River drainage and the Salmon River drainage in east-cen-
tral Idaho within the Frank Church River of No Return
Wilderness (Habeck 1992).
Lake
Classification
57
Aquatic physical - chemical factors
58
View of pond 2 which appears as two separate ponds.
View east of Salmon Mountain Lake from the ridge trail. Note island
in the lake (lower arrow) and adjoining Pond 2 (upper arrow).
View west of Salmon Mountain Lake and ridgeline. View east of Salmon Mountain Pond 3, a shallow water
body in the RNA that was not sampled.
59
Water sedge Cliff sedge
(Carex aquatilis ) (Carex scopulorum )
From Hurd et al. From Hurd et al. 1998.
Outlet from Salmon Mountain Lake flows into Pond 2. 1998.
Zooplankton
Vegetation
60
Macroinvertebrates Salmon Mountain Lake outlet
62
Allan Mountain Ponds Allan Lake is not included in the RNA because of
heavy recreational use. Lake 1 north of Allan
Mountain drains into Twin Creek. It was not sam-
Allan Mountain Research Natural Area pled. The two ponds are located in a meadow above
Salmon-Challis National Forest Allan Lake. A wetland comprised of small intercon-
nected pools is located between the ponds.
Two ponds and a lake are located in the RNA. Fred
Rabe surveyed Pond 1 on September 24, 1999.
Some data were collected from Pond 2.
Location
Geology
63
Aquatic physical-chemical factors A series of shallow pools and channels comprise a
wetland in a high meadow adjacent to Allan
Pond 1 Mountain. This wetland separates the two ponds.
Surface area (hectares): 0.01 (0.03 acres)
Length of shoreline: (m): 40 (131 ft)
Maximum depth (m): 1.2
Elevation (m): 2500 (8200 ft)
Aspect: NE
Percent shallow littoral zone: 100
Dominant bottom substrate: soft sediment
Lake edge %: herbaceous-90, conifers-10
Alkalinity (mg/l): 8
Conductivity (micromhos): 11
pH: 6.9
Inlet: seeps
Outlet: ephemeral stream
Pond 1 above. Pond 2 below occupies a cirque. Note Wetland adjacent to the two ponds. Note subalpine
subalpine larch (Larix lyallii ) in foreground. larch (Larix lyallii ). Allan Mountain at an elevation of
9154 ft is in background.
Vegetation
64
Vertebrates
Zooplankton
Pond 1 .
Diaptomus sp.
Daphnia sp.
Simocephalus sp.
Macroinvertebrates
Ponds 1 and 2
Ephemeroptera Hemiptera
Caenis sp. Limnogonus sp.)
Coleoptera Gerris sp. Spotted frog (Rana pretiosa ) Photo credit: Corkran and
Coptotomus sp. Diptera Thoms (1996).
Pelecypoda Family Chironomidae
Pisidium sp. Class Oligochaeta
65
Literature Cited
66
Fish Lake
Location
Classification
67
wetland
20
68
Zooplankton
Copepoda
Diaptomus lintoni
Orthocyclops modestus
Cyclops venustoides
Cladocera
Daphnia rosae
Chydorus sphaericus
Vegetation
69
Literature Cited
70
Square Mountain Lake Geology
Location
Classification
71
View of wet meadow east of the lake.
The inlet stream is about 0.3 meters wide and 0.5 m meters
deep. Small pea-gravel comprises the substrate. Water
temperature of the inlet was 11 degrees C.
Inlet stream to Square Mountain Lake. Note pea-
gravel substrate and herbaceous riparian growth.
72
Vegetation Inlet stream
Platyhelminthes
The entire area burned in 1919 and partially burned again in Polycelius coronata
1933. Tree cover is minimal over much of the RNA due to Trichoptera
fire and lack of adequate soil development (Wellner and Clistorina sp.
Bernatas 1980). Whitebark pine-subalpine fir (Pinus albi- Psychoglypha sp.
caulis-Abies lasiocarpa) habitat types are found in the RNA Doliphiloides sp.
together with six subalpine fir types. Douglasia idahoensis, Gramataulius sp. ?
a category two candidate and Forest Service Sensitive Plecoptera
species is found along the southwest boundary. Family Chloroperlidae
Cultus sp.
An aquatic moss (Fontinalis sp.) is commonly observed in Ephemeroptera
the outlet stream. Wetland vegetation in the RNA includes Nixe sp.
Carex nigricans, Carex scopulorum, and Juncus parryi. Ameletus sp.
Pelecypoda
Pisidium sp.
Diptera
Subfamily Tanypodinae
Macroinvertebrates
Outlet stream
Megaloptera Paraleptophlebia, a
Sialis sp. mayfly from outlet
Ephemeroptera stream. Sketch credit:
Paraleptophlebia debilis McCafferty (1983).
Ameletus sp.
Baetis tricaudatus
Diptera
Prosimulium sp.
Subfamily Tanypodinae
Subfamily Chironominae
Coleoptera
Agabus sp.
Hygrotus sp.
Hemiptera Anterior of Polycelius
Corixidae coronata a planarian flat-
worm dominant in the inlet
Trichoptera stream. Note numerous
Clistorina sp. simple eye spots. Sketch
Limnephilus sp. credit: Kolasa (1991).
73
Literature Cited
74
Dome Lake
Location
Classification
Vegetation
Vertebrates
Dome Lake is a moraine-dammed water body
at a relatively low elevation in central Idaho. Apopulation of stunted Eastern brook trout (Salvelinus fonti-
nalis) occurrs in the lake. Successful spawning takes place
because different age classes of fish were observed.
76
Belvidere Lakes and Ponds
Geology
Belvidere Creek Research Natural Area
The bedrock varies from moderately hard, well fractured
Payette National Forest granite of the Pinnacles adjacent to Lake 3 to weathered
volcanic rocks near Lake 7. The lands are characterized as
Twelve lakes and ponds exist in the RNA. From July 18-27, being geologically young and erosion is quite active (Rabe
1978 eight of these sites together with associated inlet and et al. 1979). Much of the area has rock debris from talus,
outlet streams were studied by Fred Rabe and his Aquatic loose rubble from avalanche slopes or exposed bedrock on
Biology class from the University of Idaho. the surface.
1.
5b
3
5a
4
East end of Belvidere Creek drainage showing six of the headwa- Lower Belvidere Creek in “U” shaped valley.
ter lakes and ponds studied. USGS Quads: Edwardsburg, Profile
Gap. T20N, R9E.
77
Belvidere Lakes Study
Classification
Lake 4
Pond 5a
Pond 6
Location of three lakes and five ponds studied among the twelve
water bodies in the Belvidere Creek Research Natural Area. • Subalpine, small, shallow, cirque pond
• Medium production potential
• Circumneutral water in granitic basin
• Inlet: 1 stream; Outlet: 1 stream
Classification Lake 7
Pond 1
• Subalpine, small, deep, cirque lake
• Subalpine, small, shallow, cirque pond • Medium-high production potential
• Low-medium production potential • Circumneutral water in granitic basin
• Circumneutral water in granitic basin • Inlet: 2 seeps; Outlet: 1 stream
• Inlet: none; Outlet: 1 stream
Pond 2
Lake 3
78
Aquatic physical - chemical factors Pond 2
Area (hectares): 0.6 (1.5 acres)
Pond 1 Length of shoreline (m): 341
Area (hectares): 0.4 (1.1 acres) Maximum depth (m): 1.2 (4 ft)
Length of shoreline (m): 241 (791 ft) Elevation (m): 2348 (7700 ft)
Maximum depth (m): 2.1 (7 ft) Aspect: N
Elevation (m): 2598 (8520 ft) Percent shallow littoral zone: 100
Aspect: E Dominant substrate: rubble and gravel
Percent shallow littoral zone:100 Shoreline development: 1.204
Dominant substrate: rubble Alkalinity (mg/l): 19
Shoreline development: 1.087 pH: 6.8
Alkalinity (mg/l): 10 Inlets: 3 streams
pH: 7.0 Outlets: 1 stream
Inlets: none
Outlets: 1 stream
Belvidere Pond 1. Note banding of the lake sedi- Belvidere Pond 2. The Pinnacles at an elevation of
ments. 2827 m (9273 ft) in background.
79
Lake 3 Lake 4
Area (hectares): 0.8 (2.1 acres) Area (hectares): 1.3 (3.2 acres)
Length of shoreline (m): 458 (1503 ft) Length of shoreline (m) 457
Maximum depth (m): 3.7 (12 ft) Maximum depth (m): 6.1 (20 ft)
Elevation (m): 2512 (8240 ft) Elevation (m): 2463 (8080 ft )
Aspect: NE Aspect: N
Percent shallow littoral zone: 40 Percent shallow littoral zone: 60
Dominant substrate: cobble Dominant substrate: cobble
Shoreline development: 1.411 Shoreline development: 1.103
Alkalinity (mg/l): 20 Alkalinity (mg/l): 12
pH: 7.1 pH: 7.0
Inlets: none Inlets: 1 seep
Outlets: 1 stream Outlets: 2 streams
80
Pond 5a Pond 6
Area (hectares): 0.4 (1 acre) Area (hectares): 0.6 (1.5 acres)
Length of shoreline (m): 269 (883 ft) Length of shoreline (m): 323 (1060 ft)
Maximum depth (m): 3 (10 ft) Maximum depth (m): 3.3 (11 ft)
Elevation (m): 2402 (7880 ft) Elevation (m): 2439 (8000 ft)
Aspect: N Aspect: ESE
Percent shallow littoral zone: 90 Percent shallow littoral zone:70
Dominant substrate: cobble Dominant substrate: cobble
Shoreline development: 1.208 Shoreline development: 1.184
Alkalinity (mg/l): 16 Alkalinity (mg/l): 18
pH: 6.8 pH: 6.8
Inlets: none Inlets: 1 stream
Outlets: 1 stream Outlet: 1 stream
Pond 5b
Area (hectares): 0.5 (1.3 acres)
Maximum depth (m): 3 (10 ft)
Elevation (m): 2402 (7880 ft)
Aspect: N Map of Pond 6 in the Belvidere Creek RNA.
Percent shallow littoral zone: 50
Dominant substrate: cobble
Shoreline development: 1.1
Alkalinity (mg/l): 16
pH: 6.8
Inlets: 2 streams
Outlet: 1 stream
81
Lake 7 Vegetation
Area (hectares): 0.8 (2 acres)
Length of shoreline (m): 567 (1860 ft) The majority of terrestrial vegetation consisted of subalpine fir
Maximum depth (m): 5.1 (17 feet) (Abies lasiocarpa) habitat types. Most of the understory near
Elevation (m): 2439 (8000 ft) the lakes was grouse whortleberry (Ledum glandulosum).
Aspect: S
Percent shallow littoral zone: 60 Zooplankton
Dominant substrate: cobble
Shoreline development: 1.800 Zooplankton density and richness was quite low in most
Alkalinity (mg/l): 12 lakes and ponds. Only one taxa was present in Ponds 1, 5a,
pH: 6.8 and 5b and Lakes 2 and 4, two taxa in Pond 6, and four taxa
Inlets: 2 seeps in Lake 7. In Pond 1 and Lake 3, where fish were absent,
Outlet: 1 stream large red calanoid copepods (Diaptomus sp.) were present.
Depth of the ponds may have had some effect on numbers
and kinds of zooplankton observed.
Copepoda
Diaptomus sp.
Cyclopoidea
Cladocera
Daphnia sp.
Chydorus sphaericus
Holopedium gibberum
Belvidere Lake 7.
82
Macroinvertebrates
Outlet streams
Taxa 1 2 3 4 5a 5b
Trichoptera
Limnephilus sp. X X
Asynarchus sp. X X
Imania sp.
Rhyacophila acropodes X X
Rhyacophila vaccua X
Ephemeroptera
The stonefly (Yoraperla brevis) was collected only from
Cinygma sp. X X
the outlet stream of Pond 5b.
Ephemerella infrequens X X X
Siphlonurus columbianus X
Paraleptophlebia memorialis X X Lakes and ponds
Centroptilum sp. X X
Plecoptera Taxa 1 2 3 5a 5b 7
Nemoura sp. X
Yoraperlis brevis X Trichoptera
Acroneuria californicus X X Chyranda centralia X
Isoperla sp. X Asynarchus sp. X X X
Family Chloroperlidae X Limnephilis sp. X
Diptera Hesperophylax sp. X
Simulium sp. X Ephemeroptera
Family Chironomidae X X Centroptilum sp. X X
Pelecypoda Siphlonurus sp. X
Family Sphaeriidae X Odonata
Platyhelminthes Aeshna sp. X
Order Tricladida X Coleoptera
Agabus sp. X X
Bidessus sp. X
Neuroptera
Sialis sp. X
Hemiptera
Graptocorixa sp. X
Gerris sp. X
Diptera
Family Chironomidae X X X
Pelecypoda
Family Sphaeriidae X
Hirudinea
Family Erpodellidae X X
Platyhelminthes
Order Tricladida X
Chloroperlidae, a stonefly family, was collected from the The first record of the caddisfly Chyranda centralis in Idaho
outlet stream of Lake 4. came from specimens collected from Pond 2 in the Belvidere
Creek RNA (Russell Biggam-personal communication).
83
Camp was set up adjacent to one of the study sites. All
In Belvidere Pond 2, Chyranda centralis was identified for the first
equipment which included a microscope and a folding table
time in Idaho. The case consists of pieces of thin bark arranged
was packed in by the students enrolled in Aquatic Biology
to form a straight tube with a prominent flange, like a seam along
(Zoology 503) at the University of Idaho.
each side (Wiggins 1996).
Vertebrates
Aquatic Biology Class Field Trip
Cutthroat trout (Salmo clarki ) were sampled from Lakes 4
and 7 and Ponds 2 and 5b. In Lake 7, fish collected by The field trip from July 18-27, 1978 to the Belvidere Lake
angling and gill nets had an average length of 278 mm (11 drainage was a requirement in Zoology 503 (Aquatic
1/8 inches) and an average weight of 229 gm (1/2 lb). Fish Biology) which at the time was offered at the University of
samples from the same length class in Lake 4 averaged Idaho. Support for the course was supplied by the Office of
considerably less in length and weight. Many of the fish Continuing Education at the university. The objective of the
from Lake 4 were well developed sexually compared to trout class was to acquaint students with methods of sampling
from Lake 7 that were immature. aquatic high lakes and analyzing the resulting data. An
additional objective was to provide data to justify selection
A limited number of fish were observed in Ponds 2 and 5b. of this site as a Research Natural Area.
They were considerably smaller than in Lakes 4 and 7.
Trout from Pond 2 had ready access to spawning from inlet Literature Cited
and outlet streams. However, no young of the year fish were
noted in any of the lakes, suggesting that spawning was not Biggam, R. C. (Personal communication). January 10,
successful. 1978. Moscow, ID: University of Idaho, Biological Sciences
Department.
It is reasonable to believe that a high degree of fish winterkill
exists in the water bodies since they are so shallow and Lichthardt, J.; Rust, S. 1994. Establishment record for
some of them retain ice and snow cover quite late into the Belvidere Creek Research Natural Area within Payette
summer. Lack of fish in Ponds 1, 5a, 6 and Lake 3 even National Forest. U.S. Department of Agriculture, Forest
though they were all planted at the same time in the early Service, Unpublished report on file at Intermountain Region,
1970s supported this theory. Ogden, UT. 25 p.
Westslope cutthroat (Salmo clarki). Drawing: Wiggins, G. B. 1996. Larvae of the North American caddis-
Simpson and Wallace 1978. fly genera. 2nd Edition. Toronto: University of Toronto
Press. 267 p.
84
Mystery Lake
Fred Rabe and Craig Rabe surveyed Mystery Lake and its
outlet on August 21,1998. A lower lake and two ponds in
the RNA were not surveyed.
Location
Ecoregion Section: IDAHO BATHOLITH (M332A), Lemhi Rock glaciers are moving out into the water below the head-
County; USGS Quad: MOUNT JORDAN wall of The General , a massif ridge partly encircling the lake.
Pond 2
Pond 1
Classification
Upper Mystery Lake
Lower
Lake
• Alpine, large, deep, cirque lake
• Low production potential
• Circumneutral water in a granitic basin
• Inlet: 1 stream; Outlet: 1 stream
85
A view of rock glaciers moving into the lake. Massif in back -
ground is The General. Whitebark pine (Pinus albicaulis ) in
the foreground.
86
View of substrate in littoral zone along east side of
Mystery Lake. Lower Mystery Lake.
Vegetation
Lower Mystery Lake at 2622 m (8600 ft). Pond 1 drains Whitebark Pine (Pinus albicaulis) on slope surrounding
into the lake. Mystery Lake.
87
Golden trout (Salmo aguabonita )
Cladocera
Simocephalus sp.
Copepods
Diaptomus sp.
Cyclopoida
Macroinvertebrates
Vertebrates
Location
89
Aquatic physical-chemical factors reed grass (Calamagrostis canadensis) in one
place. On the northwest side of the lake, a drier
Area (hectares) 1 (2.5 acres) and slightly more raised meadow exists. Sedges
Length of shoreline (m): 389 (1276 ft) occur here but not Carex utricularia. Labrador tea
Maximum depth (m): 3 (10 ft) (Ledum glandulosum) together with small
Elevation (m): 2335 (7660 ft) Engelman spruce (Picea engelmannii) were
Aspect: NW observed. Scattered patches of water sedge
Percent shallow littoral zone: 90 (Carex aquatilis) were noted along the lake shore.
Dominant bottom substrate: soft sediment
Shoreline development: 1.051 A species of pondweed (Potomogeton) was sub-
Lake edge %: herbs-65, shrubs, trees-15, rocks-15 merged in the water together with the blue green
Alkalinity (mg/l): 15 algae (Nostoc). A “green island” floating in the lake
Conductivity (micromhos/cm): 20 was bur-reed (Sparganium angustifolium).
pH: 7.6
Inlet: 3 seeps A willow (Salix drummondiana) was observed lin-
Outlet: 1 stream ing the outlet area close to a small patch of alder
(Alnus sinuata). Collection and identification of
plants were made by Michael Mancuso from the
Conservation Data Center in Boise, Idaho.
Outlet
90
Macroinvertebrates
Inlet stream
Trichoptera
Ecclisomyia sp.
Diptera
Procladius sp.
Lake
Ephemeroptera
Baetis tricaudatus
Paraleptophlebia sp.
Trichoptera
Lenarchus sp.
Limnephilus sp.
Diptera
Procladius sp.
Subfamily Chironominae
Subfamily Orthocladiinae
Neuroptera
Sialis sp
Coleoptera
Neoscutopterus sp.
Beaked sedge (Carex utric- Pelecypoda
u l a ri a) occurred in an
Pisidium sp.
almost pure stand in the
wet meadow on the south
Oligochaeta
side of the lake. From Hurd Family Lumbriculidae
et al. 1998. Aeshnidae
Aeshna sp.
Zooplankton
Cladocera
Graptoleberis testunaria
Chydorus sp.
Alona sp.
Vertebrates
Graptoleberis testunaria. Sketch
credit: Melanie Abell. Cutthroat trout (Salmo clarki) up to nine inches in
length were observed in the lake.
91
Literature Cited
92
Cache Creek Lakes and Pond
On August 22, 1998 Fred Rabe and Craig Rabe sampled Lake
1 and Pond 1 together with associated inlet and outlet streams
in the RNA. Lake 2 and Ponds 2 and 3 were not sampled.
Location
Ecoregion Section: CHALLIS VOLCANICS (M332F) Lemhi FS Road 086. Sleeping Deer Mountain in background at an
County;USGS Quad: SLEEPING DEER MOUNTAIN elevation of 3012 m (9881 ft).
Pond 3
Lake 1
Lake 2
Aquatic physical - chemical factors
6
6
94
View from east shore of Cache Creek Lake 1.
14
12
8
Contour depths
are in feet
N
cascade
95
The inlet stream into the lake was 11 degrees C. The substrate
is comprised of small, light colored gravel mixed with some rub-
ble-size stones. Arelatively small amount of coarse particulate
matter (CPOM) is present. Riparian vegetation is primarily
sedges. The stream which averages about 0.5 m wide and only
3 cm deep with pools up to 10 cm in depth, cascades down a
steep slope and then flows about 30 m into the lake.
Inlet stream to Cache Creek Lake 1. Note the small size rock
substrate and clarity of water.
Vegetation
96
Quillwort - Isoetes sp.
Macroinvertebrates
Zooplankton
97
Literature Cited
Cache Creek Lake 1 - inlet
Ephemeroptera McCafferty, W. P. 1981. Aquatic entomology. Boston: Jones
Cinygmula sp. and Bartlett Publishers. 448 p.
Plecoptera
Yoraperla brevis Merritt, R. W.; Cummins, K. W. 1996. Aquatic insects of
Visoka sp. North America. Dubuque: Kendall Hunt Publishers. 862 p.
Family Chloroperlidae
Trichoptera Needham, J. G.; Needham, P. R. 1962. A guide to the study
Clistorina sp. of freshwater biology. San Franciso: Holden-Day
Allomyia sp. Publishers. 108 p.
Hesperophylax sp.
Diptera Ross, C. P. 1934. Geology and ore deposits of the Casto
Family Chironomidae Quadrangle, Idaho. U.S. Geological Survey. B-854. 132 p.
Yoraperla brevis is a roachlike stonefly Sleeping Deer Mountain on right. Abandoned lookout
found in cold clean water in an erosion- tower on summit.
al habitat. (Merritt and Cummins 1996).
Sketch Credit: McCafferty 1981.
Vertebrates
98
Master Sergeant Lake and Ponds
Classification
Soldier Lakes Research Natural Area Master Sergeant Lake
Salmon-Challis National Forest
• Subalpine, small, deep, cirque-scour lake
During July 1980, Charles Wellner, Nancy Savage and Fred • Medium-high production potential
Rabe sampled Master Sergeant Lake and three ponds com- • Circumneutral water in granitic-quartz monzonite
prising the RNA. • Inlet: 1 stream; Outlet: 1 stream
Tech
Pond Sergeant
1 Pond
Master
Sergeant
Lake
Pond 2
Geology
100
Vegetation
Invertebrates
101
Vertebrates
Literature Cited
102
102
103
Little Granite Creek Lakes
(Wellner 1979). The natural area spans elevations from abou
427 m (1400 feet)where Little Granite Creek flows into the
Little Granite Creek Research Natural Area Snake River to 2863 m (9393 ft) at the summit of one of the
Nezperce National Forest peaks. The proposed RNA will contain the entire drainage o
Little Granite Creek except for some recreational exclusions.
There are five lakes and five ponds in the proposed RNA.
Norm Howse surveyed Echo Lake, Quad Lake, and He Devil
Lake on August 7-11, 1967 (Howse 1967). Fred Rabe and
Nancy Savage subsequently made observations of Baldy
Lake and Ponds 1-3 on September 27-29, 1974. Triangle
Lake was not sampled.
Location
From Riggins, Idaho, drive south about two miles to the Seven
Devils Road (FR 517) and travel to Heaven’s Gate and the
Seven Devil’s Guard Station. By trail, Quad Lake, He Devil
Lake and Echo Lake are about 9 miles from the guard Station. View west of Seven Devils Mountain Range
Classification
He Devil Lake
105
Aquatic physical - chemical factors
Echo Lake, Quad Lake and He Devil Lake were surveyed
by Norm Howse August 7-11,1967.
Echo Lake
Area (hectares): 3.5 (8.6 acres)
Length of shoreline (m): 827 (2713 ft)
Maximum depth (m): 12.2 m (40 ft)
Elevation (m): 2208 (7243 ft)
Aspect: NW
Percent shallow littoral zone: 20
Dominant substrate: silt covered rock bottom
Shoreline development: 1.3
Lake edge %: conifers-80, herbacous-10, rock-10
Alkalinity (mg/l) 17
pH: 6.5
Inlets: 2 seeps
Outlets: 1 stream
106
Quad Lake Baldy Lakes
Area (hectares1.3 (3.2 acres)
Length of shoreline (m): 549 (1801 ft) These water bodies are reached from Seven Devils Guard
Maximum depth (m): estimate, >7.6 (>25 ft) Station Campground, a distance of about 12 miles by trail.
Elevation (m): 2195 (7200 ft) A brief description follows. Detailed measurements and
Aspect: NW samples were not taken.
Percent shallow littoral zone: 20
Dominant bottom substrate: silt covered boulders Baldy Lake , square in shape, is surrounded by a young for-
Shoreline development: 1.3 est growing over the site of a fire in 1945. An extensive lit-
Lake edge %: slide rock-90, conifers-10 toral zone exists except for the southern shore which has a
Inlets: none steep drop off. The substrate is rocky and there is no inlet
Outlet: intermittent visible. The outlet is dry except for a small spring.
No aquatic vegetation was observed in Quad Lake. Baldy Pond 1 has an estimated depth of about 3 m (10 ft).
Considerable amounts of spawning gravel were observed The substrate consists of soft bottom sediments.
around most of the water body. The pond below the lake is
about 2 m deep. Baldy Pond 2 has an estimated maximum depth of 2.4 m
(8 feet). The water body has a boulder substrate and the
He Devil Lake shoreline is predominantly rocky. The lake at its widest
Area (hectares): 2.1 (5.2 acres) point is estimated to be about 37 m (120 ft).
Length of shoreline (m): 593 (1946 ft)
Maximum depth (m): estimate, 7.6 (25 ft) Baldy Pond 3 has an estimated maximum depth of about 1
Elevation (m): 2268 (7440 ft) m (3 ft). Herbaceous plant growth covers the muddy sub-
Percent shallow littoral zone: estimate, 35 strate with filamentous algae attached to the plants. No inlet
Dominant bottom substrate: rock and sediment or outlet is noted. The diameter of the pond is estimated to
Shoreline development: 1.2 be 15 m (50 ft).
Lake edge %: conifers-30, herbaceous-20, rock-50
Inlets: none According to the topographic map the main body of Little
Outlets: intermittent stream to small pond Granite Creek begins as the outlet stream of Baldy Lake but
such is not the case. Little Granite Creek probably origi-
nates as a spring draining water out of Baldy Lake through
underground channels. This spring is located somewhere
north of the side trail to Baldy Lake and south of the small
trickle that constitutes the direct overflow out to the water
bodies below.
2
3
Baldy Pond 1 with Baldy Pond 2 to the right and Baldy Pond
He Devil Lake. Note the island in the lake.
3 to the left.
107
Vegetation Macroinvertebrates
Literature Cited
Zooplankton
Echo Lake
Daphnia longispina
Diaptomus sp.
Ostracoda
Kellicottia sp.
108
Goat Lake
The RNA has one lake. The site was visited in the
1980s by Charles Wellner and Bob Moseley to
establish an RNA. Aerial photos were taken of the
area in September1999. The RNA is located in the
western Salmon River Mountains about 8.5 air
miles southeast of Riggins, Idaho (Wellner and
Tisdale 1985).
109
Literature Cited
110
Lava Butte Lake and Ponds
Classification
Lava Butte Research Natural Area
Payette National Forest Lava Butte Lake
• Subalpine, large, deep, cirque lake
• Low production potential
On July 16-17, 1998, Fred Rabe and Nancy Abbott sampled • Circumneutral water in granite-basalt basin
the lake in the RNA. Two ponds outside the RNA but near • Inlet: none; Outlet: riffle-pool type stream
the lake were also sampled.
Lava Butte Pond SW
Location • Subalpine, small, shallow, cirque pond
• Low-medium production potential
The RNAis situated in the western Salmon River Mountains • Circumneutral water in granite-basalt basin
about 25 air miles north of McCall, Idaho. • Inlet: none; Outlet: none
SW Pond
SE
Pond
Geology
111
Lava Butte Pond SW
Area (hectares): 0.6 (1.6 acres)
Length of shoreline (m): 320 (1050 ft)
Maximum depth (m): 1.8 (5.9 ft)
Elevation (m): 2317 (7600 ft)
Aspect: E
Percent shallow littoral zone: 100
Dominant substrate: boulders and soft sediments
Shoreline development: 1.097
Lake edge %: talus-65, herbaceous-35
Alkalinity (mg/l): 11
Conductivity (micromhos/cm): 30
pH: 7.3
Inlets: None
Outlets: None
Lava Butte Pond SW. Boulders near the distant shore are-
Lava Butte Pond SE. Color of lake indicates suspended
heavily mineralized. Large fairy shrimp (Branchianecta col-
fine particles in water or “rock flour.”
oradensis ) are dominant invertebrates in the water.
112
Vegetation Zooplankton
Extensive sedge beds primarily water sedge (Carex Diaptomus shoshone is a large size calanoid
aquatilis) surround Lava Butte Pond SE. copepod which is found predominantly in water
bodies without fish populations as was the case
in the southwest pond.
Macroinvertebrates
113
Stoneflies, mayflies and caddisflies were well represented
in the first order stream outlet from the lake. Lava Butte Pond SE
Ephemeroptera
Ephemerella sp.
Diptera
Ablabesmyia sp.
Family Ceratopogonidae
Family Culicidae
Family Orthocladiinae
Odonata
Ischnura sp.
Oligochaeta
Family Lumbriculidae
Pelecypoda
Pisidium sp.
Hirudinea
Helobdella stagnalis
Literature Cited
114
Steamboat Lake Geology
From McCall follow the Warren Wagon Road (FS 21) north
along the west side of Payette Lake and the North Fork
Payette River to the town of Warren a distance of about 50
miles. Continue southeast on FS Road 340 about 1.5 miles
past Warren.Turn right (southwest) onto FS Road 359 and
follow it for nine miles.Turn right again onto FS Road 361
which ends within the RNA (Savage and Wellner 1979).
115
Aquatic physical - chemical factors Vegetation
Area (hectares): 2.7 (6.7 acres) Steamboat Lake, located in a shallow drainage basin, is sur-
Length of shoreline (m): 890 (2920 ft) rounded by a forest of mostly Engelman spruce (Picea engel-
Maximum depth (m): 1.5 (5 ft) manii), lodgepole pine (Pinus contorta) and subalpine fir
Elevation (m): 2278 (7472 ft) (Abies lasiocarpa). The lake is encircled by an open fen dom-
Aspect: NE inated by water sedge (Carex aquatilis) and firethread sedge
Percent shallow littoral zone: 100 (Carex prionophylla). The yellow water lily (Nuphar polysepa-
Dominant bottom substrate: soft sediment lum) is the only aquatic macrophyte present and is distributed
Lake edge %: herbaceous-100 % somewhat uniformly throughout the lake.
Alkalinity (mg/l): 30
pH: 7.1 Other emergents surrounding the lake are small headed
Conductivity (micromhos/cm): 10 sedge (Carex illota), early sedge (C. praeceptorum), spike
Inlets: none rush (Eleocharis pauciflora), and rush (Juncus
Outlets: 1 stream mertenisanus).
Literature cited
116
Needles Lake and Pond
Ecoregion Section: IDAHO BATHOLITH (M332A), Stump Lake. Plants in foreground are Shooting
Valley County; USGS Quad: GOLD FORK ROCK Star (Dodecatheon jeffe ry i) (Lichthardt 1993).
and BLACKMARE. Photo credit: Michael Mancuso
Pond 1.
117
Literature Cited
118
Fiddle Lake Geology
119
The inlet averages 38 cm wide and 3 cm deep. The sub-
strate is small pea-gravel. The gradient is 1 percent. No
pools are present. The water temperature is 14 degrees C.
Shrubs and sedges comprise the riparian vegetation.
The outlet stream below the thin neck of Fiddle Lake aver-
ages 1.5 m wide and 10 cm deep. It is a cascade-pool type
stream characterized by a boulder substrate and 5-6 per-
cent gradient. The water temperature is 17 degrees C. The
dominant riparian growth is herbaceous vegetation.
Vegetation
120
Outlet stream from Fiddle Lake
Trichoptera
Rhyacophyla - Albertae group
Eocosmoecus sp.
Platyhelminthes
Polycelius coronata
Diptera
Corynorena sp.
Tventenia sp.
Prosimulium sp.
Nanocladius sp.
Pentaneura sp.
Plecoptera
Cultus sp.
Isoperla sp.
Pelecypoda
Pisidium sp.
The inlet stream into Fiddle Lake. Note the small size
pea-gravel comprising the substrate. Sedges and shrubs
are the dominant riparian vegetation.
Macroinvertebrates
Trichoptera
Rhyacophyla brunea Literature Cited
Plecoptera
Visoka sp. Bennett, E. H. 1980. Reconnaissance geology and geochem-
Ephemeroptera istry of the Trinity Mountain-Steel Mountain Area, Idaho.
Ameletus sp. Report 80-11. Moscow, ID: University of Idaho, Bureau of
Diptera Mines and Geology. 56 p.
Diamesa sp.
Platyhelminthes Kolasa, J. 1991. In: Ecology and classification of North American
Polcelius coronata
freshwater invertebrates. Boston: Academic Press. p. 158.
121
Lichthardt, J. 1993. Establishment record for Trinity
Mountain Research Natural Area within Boise National
Forest. U.S. Department of Agriculture, Forest Service,
Unpublished report on file at Intermountain Region, Ogden,
UT. 20 p.
122
123
124
Surprise Valley Lake and Pond
Location
Lower pond
Classification
125
Aquatic physical - chemical factors Upper Surprise Valley Lake
126
Inlet stream into Upper Surprise Valley Lake.
Temperature was 9 degrees C. First order tributary (riffle-pool stream type) of Fall Creek from
upper lake. Standhope Peak in the background.
Vegetation
127
Inlet to Upper Surprise Valley Lake
Diptera
Diamesa sp.
Prosimulium sp.
Trichoptera
Lenarchus sp.
Zooplankton
128
Smiley Mountain Lake and Ponds Geology
Location
Classification
129
Percent shallow littoral zone: 100
Dominant bottom substrate: soft sediments, silt
Shoreline development: 1.059
Lake edge %: herbaceous / willow-80, talus, cliffs-20
Alkalinity (mg/l): 7
pH: 7.0
Conductivity (micromhos/cm): 15
Inlets: 1 seep
Outlets: 1 stream
The inlet stream into Lower Smiley Mountain Pond originat- View south of Lower Smiley Mountain Pond.
ed from snow and ice accumulated under loose rock. At its
beginning the inlet is about 1 m wide and 8 cm deep. It The low gradient outlet from the lower pond with a boulder-
flows rapidly 100 m downhill into a shallow pool which then rubble substrate averages about 61 cm wide and 13 cm
empties into the pond. At this point, the stream is about 45 deep.The riparian vegetation is primarily sedges and other
cm deep. herbaceous growth.
131
A large number of interconnected ponds within an extensive
wet meadow occur below the study site.
Vegetation
132
Outlet Upper Smiley Mountain Lake
Ephemeroptera
Baetis bicaudatus - dominant
Plecoptera
Family Chloroperlidae
Diptera
Subfamily Tanypodinae
Subfamily Chironominae
Simulium sp.
Trichoptera
Lenarchus sp.
Platyhelminthes
Procotyla sp.
133
Outlet Lower Smiley Pond First order tributary of Lake Creek below Lower
Smiley Pond.
Ephemeroptera
Baetis bicaudatus Ephemeroptera
Trichoptera Baetis bicaudatus
Dicosmoecus sp. Ameletus sp.
Diptera Plecoptera
Simulium sp. Zapada sp.
Subfamily Orthocladiinae Trichoptera
Platyhelminthes Chyranda sp.
Procotyla sp. Dicosmoecus sp.
Psychoglypha sp.
Diptera
Inlet Lower Smiley Pond Pseudodiamesa sp.
Ephemeroptera Simulium sp.
Baetis bicaudatus Triboles sp.
Trichoptera Pagastea sp.
Rhycophyla albertae Diamesa sp.
Diptera Platyhelminthes
Subfamily Orchocladinae Proctyla sp.
134
Second order tributary of Lake Creek below conflu-
ence of stream draining Lower and Middle Smiley
Mountain Ponds
Ephemeroptera
Baetis bicaudatus
Cinygmula sp.
Seratella tibialis
Epeorus albertae
Trichoptera
Rhyacophyla albertae
Rhyacophyla bettini
Dicosmoecus sp.
Parapsyche sp.
Arctopsyche sp.
Diptera
Simulium sp.
A freshwater shrimp (Gammarus lacustrus ) occurs in Upper
Smiley Mountain Lake.and Middle Smiley Pond. This crus-
tacean is usually not found in high lakes where fish occur as
was the case with the Smiley lakes. Gammarus is omniver -
ous. It browses on the film of diatoms and organic debris cov-
ering leaves such as might occur on the dense sedges along
the lake shores. Hyallela azteca is another freshwater shrimp
sampled from the two lakes. It is considerably smaller in size
than Gammarus.
Vertebrates
135
Literature Cited
136
137
138
Kenney Creek Pond
Pond
View into Kenney Creek drainage (arrow) and the
Continental Divide separating Idaho from Montana.
139
Literature cited
140
Upper Mill Lake Geology
The larger Mill Lake is a recreational site and not part of the
Mill Lake Research Natural Area RNA. Subalpine and alpine habitats above Mill Lake comprise
Salmon-Challis National Forest the RNA. Upper Mill Lake is situated in a cirque at timberline.
Several shallow ponds exist in the basin just below Upper Mill
Charles Wellner, Nancy Savage and Fred Rabe visited the Lake. The lake basin consists primarily of a quartzite forma-
RNA on November 3, 1975 and sampled the lake, three tion. Steep cliffs below the crest and numerous moraines
ponds and outlet streams. A fourth pond was dry at the time down-valley are evidence of alpine glaciation (Wellner 1993).
of the survey.
Location Classification
Upper Mill Lake
Mill Lake is located on the east slope of the Lemhi Range • Alpine/subalpine, small, deep, cirque lake
about 15 miles west-southwest of Leadore, Idaho. • Circumneutral water in quartzite basin
• Low production potential
Ecoregion Section: BEAVERHEAD MOUNTAINS (M332E), • Inlet: seeps; Outlet: meandering glide stream
Lemhi County; USGS Quad: MOGG MOUNTAIN
1 2
Lake
View east of Upper Mill Lake at 2863 m ( 9390 ft) elevation. Part
of the lake is above timberline. Three lower ponds lie east of the
Pond lake. The water bodies are in a “U” shaped glaciated valley.
1 Pond 3
Pond 2 The outlet of Upper Mill Lake flows through talus rubble and
reappears as a slow moving low gradient stream in a sphag-
Upper Mill num-sedge meadow. The stream substrate is comprised of
Lake rubble, gravel and sediment. The outlet averages about 1 m
wide and 8 cm deep and flows about 200 m to Pond 1. The
moss Dermatocarpo fluviatile covers rocks in the stream.
141
The outlet from Pond 1 discharges to Pond 2; the Pond 2
outlet discharges to Pond 3. The water bodies all appear to
be less than two m deep. Pond 4 was dry in November.
Upper Mill Lake looking southwest into talus slope. Note lit-
toral zone in foreground. Pond 1 receives water from Upper Mill Lake.
142
Filamentous green algae (Zygnema
sp.) Sketch credit: Palmer (1959).
Zooplankton
Macroinvertebrates
Ephemeroptera Ephemeroptera
Centroptilum sp. Centroptilum sp.
Trichoptera Trichoptera
Family Limnephilidae Brachycentrus sp.
Diptera Diptera
Dolichopesa sp. Family Chironomidae
Coleoptera
Agabus sp.
Vegetation
143
Literature Cited
144
Upper Merriam Lake
Location
Geology
Area (hectares): 0.6 (1.5 acres) Upper Merriam alpine lake is located at 3122 m (10,240 ft) eleva-
Length of shoreline (m): 459 (1506 ft) tion. The lake lies against a cirque headwall surrounded by talus
Maximum depth (m): estimate > 5 (16 ft) rock on three sides. One year it was reported that lake size varied
Elevation (m): 3122 (10,240 ft) from about 1.6 h (4 acres) during snowmelt to less than 0.24 h (1
acre) after snow melt. The water is clear and the bottom substrate
Aspect: SE
consists of large angular rocks and silt.
Percent shallow littoral zone: estimate 20
Dominant substrate: boulders and silt
Shoreline development: 1.471
Lake edge %: talus rock-100
Alkalinity (mg/l): 41-105 recorded in lake and pools
Inlet: small seeps
Outlet: riffle-pool stream
146
The lake outlet and other steep first order streams enter a wet
meadow downstream where the gradient diminishes and the
stream becomes sinuous with fine organic and inorganic sub-
strate materials. The stream eventually flows into lower
Merriam Lake. A portion of the meadow is of interest because
of its hummocky character probably caused by freezing and
thawing action. Seasonal pools are brown in color (dystrophic).
Vegetation
147
Macroinvertebrates
148
149
150
Mount Harrison Pond
Location
Ecoregion Section: NORTHWESTERN BASIN AND Looking down at pond (arrow) from inside Forest
Service fire lookout atop Mount Harrison.
RANGE (342B). Cassia County; USGS Quad: MOUNT
HARRISON
Geology
From Albion, Idaho travel southeast on State Route 77 for
about 4 miles to the intersection with the Howell Canyon
Mount Harrison is the highest point in the Albion Range of
Road which becomes FS Road 549. Drive about 9 miles to
southern Idaho. The steep topography and distribution of
the summit. From the Forest Service fire lookout, descend
geologic formations result in very distinct vegetation pat-
approximately 600 feet to the pond below.
terns (Mancuso and Evenden 1996). Subalpine fir (Abies
lasiocarpa) occured on quartzite while limber pine (Pinus
albicaulis) was observed on calcareous substrates. The
basin consists of a steep-walled, rocky cirque with a vernal
pool at the bottom.
Pond
Classification
151
Aquatic physical - chemical factors
Vegetation
View looking northwest at Mt. Harrison Pond. The pond is Castilleja christii (Indian paintbrush) is endemic to Mount
ephemeral drying up in some years. Surface water had Harrison and is a candidate for listing. (Mancuso and
receeded about 6 m (20 ft) by the middle of August. Snow Evenden 1996). Cymopterus davisii is endemic to the
drifts in the basin likely provide an inflow to the pond.
Albion Range and is known from only two other sites.
Machaeranthera laetevirens (aster) is a USFS Sensitive
An army plane crashed into the basin in 1942 killing a few species and known from three other sites in Idaho and
people. A large tire and other scattered debris still remain Nevada. Mount Harrison RNA is the only documented
from the crash close to the pond. occurrence for this species in Idaho.
152
Zooplankton
Macroinvertebrates
Backswimmers (Notonecta undulata and N. kirbiyi ) were
found throughout the pond but were in more dense concen-
Trichoptera trations in the vicinity of Eleocharis, a species of sedge
Clistorina sp. growing in the north end of the pond.
Diptera
Subfamily Chironominae
Hemiptera
Notonecta undulata -dominant
Notonecta kirbiyi
Family Corixidae
Coleoptera
Agabus sp.
Eubranchiopoda
Branchianecta paludosa
153
Literature Cited
154
155
156
Bloomington Lake
Location
Pond 2
Pond 1
Classification
Zooplankton
Bloomington Lake
158
Bloomington Pond 2
Diptera
Subfamily Orthocladiinae
Hemiptera
Gerris sp.
Odonata
Family Aeshnidae
Pelecypoda
Pisidium sp.
Gastropoda The whirligig beetle (Gyrinus ) is somewhat
Physella sp. unique in having a ventral pair of eyes that
Oligochaeta serve for vision in water and a dorsal pair used
Family Lumbriculidae for aerial vision, an adaptation for optimal sight
at the water surface. The beetles swim errati-
cally or dive while emitting defensive secre-
tions when disturbed. McCafferty (1983). .
Sampling the lake was difficult because of the limited littoral
zone. Density of macroinvertebrates was high possibly due
to alkaline water conditions in the basin. The lake and Pond
2 shared very few common species.
159
Literature Cited
160
Summary and Discussion
Classification elements
Four classification elements of the water bodies are summarized in Table 4. Two lakes and
two ponds are montane; 20 lakes and 16 ponds are subalpine; and five lakes and two ponds
are alpine. Alpine water bodies are located in the Broad Valleys, Sawtooths and Idaho
Batholith subregions. Montane sites are in the North Idaho, Idaho Batholith and Western
Fringe subregions. Subalpine waters are located in all subregions.
Twenty-one of the lakes are small in size or less than 4 hectares (10 acres) and six are
large or more than 4 hectares in size (Table 4). Twenty-one of the lakes are classed as
deep or more than 4.5 m (15 ft) and six lakes are shallow or less than 4.5 m in depth. All
of the ponds are small and shallow.
Ten lakes and three ponds are classed as having low production potential; 16 lakes and
16 ponds have low-medium, medium and medium-high production potential and; one lake
and one pond have a high production potential (Table 4).
Production potential is mostly low in alpine lakes. Factors that contribute to low production are
high elevation, small littoral zone, boulder-bedrock substrate and few or scattered sedge mats.
Exceptions occur in Upper Smiley Lake and Surprise Lake in the Pioneer Mountains where
extensive sedge beds occur together with relatively large littoral zones of cobble and rubble.
161
162
Montane lakes have medium-high to high production potential. Fish Lake (high produc-
tion) has a low elevation, large littoral zone with a mix of small rock and sediments com-
prising the bottom substrate, high shoreline development, extensive sedge mats and a rel-
atively high alkalinity. Subalpine lakes varied from low to medium-high production and
subalpine ponds from low to high production potential (Table 4).
In comparing subalpine lakes with forest lakes located near the base of Mount Rainier,
Larson et al. (1994), observed that lower elevation forest lakes had larger watersheds,
larger surface areas, deeper waters and more nutrients than subalpine lakes. This was
true for the two montane lakes in this study compared to most subalpine waters.
Lake origin
Thirty-four water bodies were characterized as having a cirque origin formed near a
mountain headwall. Fourteen lakes/ponds were classed as cirque-scour occupying basins
carved in less resistant rock formed down valley from the headwall. It was often difficult to
differentiate between cirque and cirque-scour types due to variation in the extent and flow
patterns of glaciers.
Paternoster lakes and ponds form a chain of at least three water bodies connected by a
stream in a glaciated valley.Two sets of paternoster lakes were observed in this study. For
example, in the Graves Peak watershed, a small pond is linked to a large lake by a steep
gradient stream merging into a cascade-pool type stream. The outlet to the lake becomes
a steep cascade that flows into a lower pond. A pond further down stream is connected
by a steep gradient stream.
The two montane lakes in the study are formed by an end moraine in extended valleys.
Upland lakes and ponds form in depressions scoured by ice caps on gently rolling or upland
valleys. This type might represent early Wisconsin ice cap glaciation or an even older pre-
Wisconsin period (Rabe and Breckenridge 1985). Three Ponds RNA is a montane site mod-
ified by the presence of beaver activities that control water levels in the ponds.
Semidrainage systems
Semidrainage waterbodies are not connected with definite drainage systems or the
drainage is poorly defined (Pennak 1989). Pennak studied semidrainage ponds in
Colorado and described most as being located in grassy meadows with highly organic,
peaty soils and no surface outlets or having poorly defined outlets. Maximum depths range
from 1 to 4 m, and water levels remain somewhat constant. Most semidrainage water-
bodies in the Idaho study conform to the above meadow description; however Lava Butte
SW is a semidrainage pond set in a rocky basin with limited organic growth.
163
Since semidrainage waters are for the most part closed systems, their basins fill with fine
organic sediment that supports rooted plants such as burr reed (Sparganium angustifoli-
um) one of the most common aquatic macrophytes observed in this study. Also amphib-
ians, leeches, aquatic earthworms, dragonflies, and snails are apt to occur in these mead-
ow waters more than in rocky drainage basins.
Theriault Pond is a small, shallow, montane waterbody having a substrate comprised of soft
organic sediments and encircled by an extensive sedge meadow. In addition a large popula-
tion of spotted frogs (Rana pretiosa) occupies this site. It is not a closed system since a rela-
tively deep outlet stream exits the pond. However further downstream the outlet diminishes in
size so that later in the year it might be described as ephemeral. Cache Creek pond has some
of the same characteristics described above but also has a well defined outlet stream.
Topographic maps are not often accurate in showing or not showing outlets.
Modifiers
All of the waterbodies in this study are circumneutral (pH 5.5-7.4) except Steep Lakes
which are alkaline (pH 7.5-8.4) and Upper Merriam Lake and Bloomington Lake which is
highly alkaline (pH > 8.4).
Medium soft water (13-30 mg/l) most commonly occurs in Precambrian Belt and volcanic
rock basins compared to soft water lakes (0-12 mg/l) mostly in granitic basins.
Bloomington Lake in a quartzite-dolomite basin is a hardwater lake with an alkalinity of
104 mg/l. Merriam Lake basin pools and lake are hardwater with alkalinities of 41-105
mg/l. They occur in limestone, dolomite, sandstone, quartzite basins. Steep Lakes in
Precambrian Belt rock are also classed as hardwater with alkalinities of 50 mg/l.
164
Surface and subsurface seepage commonly form inlets into alpine lakes such as Mystery
Lake which is close to the headwall. Such small flows are often difficult to distinguish and
usually do not appear on maps. Examples of multiple inlets into subalpine large deep
lakes were observed at Fenn Lake with four inlet streams and Fish Lake with 27 stream
inlets and nine seep inlets. A cascade of water was observed to flow into Graves Peak
Pond 2 from the lake above, part of the paternoster pattern. Outlet streams are either rif-
fle-pool, meandering glide or cascade-pool type streams. Riffle-pool and meandering
glide streams are the most common types observed.
Inlets and outlets often differ in size, substrate, temperature and biological composition.
Inlet streams are usually small in width and depth and have small size substrate, com-
monly small gravel. Inlets are usually several degrees cooler than outlet streams. Outlet
streams have more coarse particulate organic matter and dense filamentous algae often
covers the rocks late in the season. When this occurred, fewer macroinvertebrates were
present in the stream channel.
Biota
The most common aquatic plants associated with standing water are quillwort (Isoetes sp.)
and bur-reed (Sparganium) Fontinalis is a moss prevalent in streams. Twenty-three different
sedges and rushes were identified. Juncus mertenisana is the rush most frequently collect-
ed and Carex aquatilus and C. scopulorum are the most common sedges.
There are 16 different mayflies (Ephemeroptera) collected. Dominant forms are listed in
Table 5. Other common mayflies sampled are Ameletus and Nixe. Twelve different
Plecoptera (stoneflies) are recorded but only from streams since they do not ordinarily
occur in lakes. Thirty-three caddisfly (Trichoptera) taxa are identified. Dominant mayflies,
stoneflies and caddisflies are listed in Table 5.
165
Table 5. Dominant Ephemeroptera, Plecoptera and Trichoptera from lakes, ponds and streams
Insect representatives from the Orders Coleoptera, Hemiptera, Diptera, Megaloptera and
Odonata are also present. Macroinvertebrates other than insects common in the water
bodies are freshwater clams (Pisidium), freshwater shrimp (Gammarus lacustris, Hyallela
azteca), flatworms (Polycelius ), and fairy shrimp (Anostraca).
Recommendations
The classification system presents elements and modifiers (Tables 1 and 3) that can be
observed, measured and analyzed. This approach has two primary benefits. First, it sets
up uniform methods and target parameters for future research. Second, the system can
be applied to gap analysis to identify missing or under-represented natural area types.
These gaps may not exist on the landscape or may simply not be reported in the inven-
tory. Future research efforts can focus on covering the gaps and bringing more natural
areas into the RNA system.
Of the 52 lakes and ponds assessed in this study, only five were classed as alpine, and
lakes in only three RNAs were identified as highly alkaline. Our surveys have identified a
series of alpine lakes and ponds located in sedimentary rock basins adjacent to waters in
granitic basins in the White Cloud Mountains of central Idaho. Highly alkaline alpine lakes
in argillaceous quartzite, dolomite and conglomerates at elevations above 9000 feet are
common here (Wissmar and Rabe 1967). Such lakes would provide an excellent oppor-
tunity for research comparing the composition and density of macroinvertebrate and plant
communities in soft and hard water lakes.
166
There is also a need to establish more upland and moraine lakes as RNAs. Both of these
geomorphic types occur at slightly lower elevations and are not recognized as high lakes.
However since they demonstrate such significant differences in limnological characteris-
tics, a greater effort should be made to identify and inventory these systems.
High lake ecosystems provide an important early warning system for environmental prob-
lems expected to grow in number, such as the threat of acid rain and eutrophication.
Natural perturbations can also be monitored, as when ash from Mount St. Helens fell on
the Steep Lakes, where we had previously gathered baseline data (Crumb 1977).
It is important to increase public awareness of the opportunities that RNAs offer for studying
natural systems. Educators and research groups can use these RNA lakes and ponds to
give students valuable experience in making scientific observations and collecting samples.
Such learning opportunities are exciting for those who enjoy and appreciate nature and can
open up a new perspective for those who have never been introduced to these prized nat-
ural systems. At the same time, the research produces more information about high lake
ecosystems and heightens our awareness and appreciation of Idaho’s natural treasures.
Literature Cited
Crumb, S. 1977. Long term effects of fish stocking on the invertebrate communities of
Steep Lake Idaho. Moscow ID, University of Idaho. 27 p. Thesis.
Larson, G.L.; Wones, A.; McIntire, C. D.; Samora, B. 1994. Integrating limnological char-
acteristics of high mountain lakes into the landscape of a natural area. Environmental
Management 18 (6): 871-888.
Rabe, F. W.; Elzinga, C.; Breckenridge, R. 1994. Classification of meandering glide and
spring stream natural areas in Idaho. Natural Areas Journal 14 (3): 188-202.
Wissmar, R. C.; Rabe, F. W. 1967. Preliminary study of some White Cloud Mountain lakes
in central Idaho. Biological Sciences Department, University of Idaho, Moscow, ID.
167
Glossary
Terms that are in bold italics and other words describing high mountain lakes are includ-
ed in the glossary. The glossary was adapted in part from Horn and Goldman (1994),
Hutchinson (1967, Merritt and Cummins (1996, and Reid (1964).
Alkalinity— Ability of lake water to neutralize acid expressed in terms of calcium carbon-
ate as mg/l.
Belt Rock — Formations of sedimentary rocks laid down in Idaho, eastern Washington
and western Montana during the latter part of Precambrian time, 850-1450 million years
ago.
Cirque — A glacially formed depression containing a lake at the upper end of a moun-
tain valley.
Cirque-scour — Basin scoured out down-valley by alpine glaciers, usually in less resis-
tant rock.
Conductivity— Measure of water to carry an electrical current, varies both with the
number and type of ions the solution contains. It is expressed as micromhos/cm.
CPOM — Coarse particulate organic matter comprised of wood litter, leaves and nee-
dles in water bodies.
Granite — Common intrusive igneous rock compound predominantly of quartz and fel-
sic minerals with crystals large enough to distinguish with the unaided eye.
168
Hard water — Waters that contain bound carbon dioxide in excess of 30-35 mg/l.
Common in regions where the substrate contains easily dissolved minerals.
Lake — Large inland body of water. This study used a curve plotting maximum depth in
meters against area in hectares used to designate waters as lakes or ponds.
Littoral —Shallow zone. In this study lake waters less than 3 meters (9.8 feet) in depth.
Macroinvertebrate — Invertebrates visible with the unaided eye. Aquatic insects, flat-
worms, freshwater clams, freshwater shrimp, fairy shrimp and snails are examples of
macroinvertebrates collected in lakes and streams.
Modifiers— Lake characteristics (pH, rock type ,hydrology) that provide supplementary
information to classifying high mountain lakes.
Montane — a mountain zone lower than alpine and subalpine zones. Upland and moraine
type lakes are commonly found in this zone.
Moraine — A deposit of glacial till. One of the geomorphic lake types in this study char-
acterized as being formed by morainal dams.
Paternoster — Variety of cirque-scour lakes that form a linear chain of at least three lakes
connected by a stream in a glacial valley.
Pond — Small shallow body of water in a depression. A curve plotting maximum depth
in meters against area in hectares is used for designating waters as lakes or ponds.
Production potential — Production of lake based on seven physical, chemical and biot-
ic parameters. Lakes are classified as poor, medium-poor, medium, medium-high and
high production.
169
Reference area —Pristine or unaltered site used as a contro to compare with locales
changed by human disturbance or natural events.
Riffle-pool — Stream type with series of alternating stretches of fast and slow water in
the channel located on a terrain of moderate gradient.
RNA — Research Natural Area. Tracts of land and water set aside by the U.S. Forest
Service for the purpose of research, biotic diversity, education and reference areas.
Rock glacier —Mass of poorly sorted angular boulders and other material cemented by
interstitial ice, as observed in Mystery Lake.
Scree — An accumulation of rocky debris lying on a slope or at the base of a hill or cliff;
talus
Sedge — Any of various plants of the family Cyperaceae resembling grasses but having
solid stems.
Sedimentary rock — A deposit of sediment hardened into rock. Most sedimentary rocks
are layered or stratified.
Seep — Slow movement of water, sometimes derived from melting snow through porous
material.
Semidrainage lake — Small, shallow, soft bottomed lakes lacking a permanent outlet and
not being connected with a definite drainage system, most commonly located in a mead-
ow. Macroinvertebrate community is usually different compared to that in a seepage lake.
Shoreline development — Amount of shoreline relative to size of lake. The more convo-
luted or irregular the shoreline, the more “edge” or shoreline development of the lake.
Lakes with more edge or contact with shore have higher production potential than those
more round.
Soft water lake — Having a low pH. Concentratrion of bound carbon dioxide as carbon-
ate is low usually less than 10 mg/l.
Subalpine lake — Located on a high upland slope but not above tree line.
Subregion — An area clssified as similar geology, terrain, climate and plant cover. There
are ten subregions in the state of Idaho, seven of which contain high mountain lake
Research Natural Areas.
Upland lake — Form in depressions scoured by ice caps on gently rolling or upland val-
leys.
170
Volcanic rock — Finely crystalline or glassy igneous rock resulting from volcanic action
at or near the earth’s surface, either ejected explosively or extruded as lava, e.g. basalt.
Literature Cited
Horne, A.. J.; Goldman, C. R. 1994. Limnology. Second Edition. New York: McGraw Hill
Publishers. 576 p.
Reid, G. K. 1964. Ecology of inland waters and estuaries. New York: Reinhold Publishing.
373 p.
171
Appendix A: Vascular and Nonvascular Plant Species of High
Mountain Lake RNAs in Idaho
The emphasis here is on aquatic and semi-aquatic plants, however some terrestrial
species in describing an RNA are also listed. Vascular plant nomenclature follows
Hitchcock and Cromquist (1973). Common names are developed by the Northern Region,
U.S. Department of Agriculture, Forest Service (U. S. Department of Agriculture, Forest
Service 1992). Moss nomenclature follows Anderson and others (1990).
Pteridophytes
Isoetaceae
Isoetes lacustris L. Lake quillwort
Dry Opteridacea
Althyrium filix-femina (L.) Roth ex Mertens Ladyfern
Gymnosperms
Cupressaceae
Thuja plicata Donn. Western redcedar
Pinaeceae
Abies grandis (Dougl.) Forbs Grand fir
Abies lasiocarpa (Hook) Nutt. Subalpine fir
Larix lyalli Parl. Subalpine larch
Picea contorta Dougl. Lodgepole pine
Picea engelmannii Parry Engelmann spruce
Pinus albicaulis Engelm. Whitebark pine
Pinus flexilis James Limber pine
Tsuga heterophylla (Raf.) Sarg. Western hemlock
Tsuga mertensiana (Bong.) Carr. Mountain hemlock
Angiosperms
Betulaceae
Alnus sinuata (Regel) Rydb. Sitka alder
Betula papyrifera Marsh. Paper birch
Boraginaceae
Romanzoffia sitchensis Bong Mistmaiden
Cyperacea
Carex aquatalis Wahl. Water sedge
Carex canescens L. Gray sedge
Carex dioica L. Yellow-bog sedge
172
Carex geyeri Boott Elk sedge
Carex illota Bailey Small-headed sedge
Carex lasiocarpa Ehrh. Slender sedge
Carex lenticularis Michx. Lens sedge
Carex limosa L. Mud sedge
Carex mertensii Prescott Merten sedge
Carex microptera Mack. Small-winged sedge
Carex multicostata Mack. Many-ribbed sedge
Carex nigricans C. A. Meyer Black alpine sedge
Carex oderi Retz. Green sedge
Carex parryana Dewey Parry sedge
Carex prionophylla Holm Firethread sedge
Carex raynoldsii Dewey Raynold sedge
Carex rossii Boott Ross sedge
Carex scopulorum Holm Holm Rocky Mountain sedge
Carex utriculata Boott Beaked sedge
Carex vesicaria L. Inflated sedge
Eleocharis pauciflora (Lightf.) Link Few-flowered spike rush
Compositae
Arnica alpina (L.) Olin Alpine arnica
Machaeranthera laetevirens Ness Aster
Crassulaceae
Sedum borschii Stonecrop
Ericaceae
Gaultheria humifusa (Grah.) Rydb. Wintergreen
Kalmia microphylla (Hook.) Heller Laurel
Ledum glandulosum Nutt. Labrador- tea
Menziesis ferrusginea Smith Fool’s huckleberry
Rhododendron albiflorum Hook White rhododendron
Vaccinium scoparium Leiberg Grouse whortleberry
Vaccinium spp. Huckleberry
Gramineae
Deschampsia cespitosa (L.) Beauv. Hairgrass
Hippuridaceae
Myriophyllum sp. L. Water-milfoil
Hydrocharitaceae
Vallisneria americana Michx. Tapegrass
Juncaceae
Juncus drummondii E. Meyer Drummond’s rush
Juncus ensifolius Wikst. Dagger-leaf rush
Juncus filiformis L. Thread rush
173
Juncus mertensianus Bong Rush
Juncus nevadensis Wats Nevada rush
Juncus parryi Engelm. Rush
Juncus torreyi Cov. Rush
Menyanthaceae
Meyanthes trifokiata L. Bogbean
Nymphaeaceae
Nuphar polysepalum Engelm. Yellow water- lily
Umbelliferae
Cymopterus sp. Raf. Cymopterus
Musineon lineare Raf. Rydberg’s musineon
Poaceae
Callamagrostis canadensis (Michx) Beauv. Bluejoint reedgrass
Festuca viridula Vasey Green fescue
Potamogetonaceae
Potomegeton crispus L. Curly pondweed
Potomegeton natans L. Floating-leaved pondweed
Primulaceae
Douglasia idahoensis Henderson
Ranunculaceae
Calltha leptosepala DC. Marshmarigold
Saxifragaceae
Saxifraga oppositifolia L. Purple saxifrage
Salicaceae
Salix artica Pall. Arctic willow
Salix commutata Bebb Undergreen willow
Saoix drummondiana Barratt Drummond willow
Salix planifolia Pursh Planeleaf willow
Scrophulariaceae
Castilleja christii Rydb. Indian paintbrush
Sparganiaceae
Sparganium angustifolium Michx. Bur-reed
174
Mosses
Literature Cited
Anderson, L. E.; Crum, H.; Buck, W. 1990. List of mosses of North America north of
Mexico. The Bryologist 93(4): 448-499.
Hitchcock, C. L.; Cromquist, A. 1973. Flora of the Pacific Northwest. Seattle, WA:
University of Washington Press. 730 p.
175
Appendix B: Zooplankton Species of High Mountain Lake RNAs in Idaho
Cladocera
Polyphemidae
Polyphemus pediculus (Linne)
Holopedidae
Holopedium gibberum Zaddach
Chydoridae
Alona sp. Baird
Chydorus sphaericus (O. F. Muller)
Chydorus spp. Stebbing
Graptoleberis sp. Sars
Pleuroxus sp. Baird
Daphnidae
Ceriodaphnia spp. Dana
Daphnia rosea Sars
Daphnia schodleri Sars
Daphnia spp. Straus
Scapholeberis sp. Schodler
Simocephalus sp. Schodler
Bosminidae
Bosmina longirostris (O. F. Miller)
Bosmina spp. Sars
Copepoda
Diaptomidae
Diaptomus arapahoensis Dodds
Diaptomus lintoni S. A. Forbes
Diaptomus shoshone S. A. Forbes
Diaptomus spp.
176
Cyclopidae
Macrocyclops albidus Claus
Cyclops venustoides Coker
Orthocyclops modestus E. B. Forbes
Cyclopidae spp.
Harpacticoida
Ostracoda
Literature Cited
Brooks, J. L. 1963. Cladocera 1963. In: Fresh water biology. New York: Wiley Press. 1248 p.
Wilson, M. S.; Yeatman, H. C. 1963. Free-living Copepoda. In: Fresh water biology. New York:
Wiley Press. 1248 p.
177
Appendix C: Macroinvertebrate taxa of High Mountain
Lake RNAs in Idaho
Immature aquatic insects were identified from Merritt and Cummins (1996), Wiggins
(1996) and Stewart and Stark (1993). Pennak (1989) was used to identify macroinverte-
brates other than aquatic insects. The invertebrates were collected from both lakes and
streams.
ARTHROPODA
INSECTA
Ephemeroptera
Heptageniidae
Cingma sp.
Cinygmula sp.
Epeorus albertae (McDunnough)
Epeorus deceitivus (McDunnough)
Nixe sp.
Ephemerillidae
Ephemerella infrequens Eaton
Seratella tibialis McDunnough
Ameletidae
Ameletus similor Eaton
Ameletus sp.
Caenidae
Caenis sp.
Leptophlebiidae
Paraleptophlebia debilis (McDunnough)
Paraleptophlebia memorialis (McDunnough)
Siphlonuridae
Siphlonurus columbianus
Siphlonurus sp.
Baetidae
Baetis bicaudatus Dodds
Baetis tricaudatus Dodds
Callibaetis sp.
178
Centroptilum sp.
Plecoptera
Peltoperlidae
Yoraperla brevis (Banks)
Nemouridae
Amphinemura sp.
Nemoura sp.
Visoka sp.
Zapada sp.
Perlidae
Acroneuria sp.
Perlodidae
Cultus sp.
Isoperla sp.
Setvena bradleyi (Smith)
Chloroperlidae
Alloperla sp.
Sweltsa sp.
Trichoptera
Glossosomatidae
Glossosoma sp.
Protoptila sp.
Limnephilidae
Asynarchuis sp.
Chyranda sp.
Clistorina sp.
Desmona sp.
Ecclisocosmoecus sp.
Ecclisomyia sp.
Ecosmoecus sp.
Grammotaulius sp.
Hesperophylax sp.
Homophylax sp.
Lenarchus sp.
Limnephilus sp.
179
Onocosmoecus sp.
Hydropsychidae
Arctopsyche sp.
Parapsyche sp.
Philopotamidae
Dolophilodes sp.
Polycentropoidae
Polycentropus sp.
Apataniidae
Allomyia sp.
Brachycentridae
Brachycentrus sp.
Uenoidae
Neothremma sp.
Odontoceridae
Megaloptera
Sialidae
Sialis sp.
Coleoptera
Dytiscidae
Agabus sp.
Coptoptomus sp.
Hydroporus sp.
Hydrovatus sp.
Hygrotus sp.
Ilybius / Agabus sp.
Neoscutopteris sp.
Rhantus sp.
Uvarus sp.
Gyrinidae
Gyrinus sp.
Limnogonus sp.
180
Hemiptera
Notonectidae
Notonecta kirbyi Hungerford
Notoecta undulata Say
Corixidae
Diptera
Ceratopogonidae
Bezzia sp.
Stratiomyidae
Stratiomyia sp.
Culicidae
Psychodidae
Simulidae
Prosimulium sp.
Chironomidae
Ablabesmyia sp.
Chironominae
Corynorena sp.
Cryptolabis sp.
Diamesa sp.
Dolichopesa sp.
Hydrobaenus sp.
Microspectra sp.
Nanocladius sp.
Orthocladiinae
Pagastea sp.
Pentaneura sp.
Polypedilum sp.
Procladius sp.
Pseudodiamesa sp.
Tanypodinae
Tventenia sp.
Odonata
Cordulegastridae
181
Libelluidae
Aeshnidae
Aeshna sp.
Coenagrionidae
Ischnura sp.
Ischnura / Enallagma
CRUSTACEA
Gammaridae
Gammarus lacustris Sars
Talitridae
Hyallela azteca Saussure
Branchinectidae
Branchinecta coloradensis Packard
Branchinecta paludosa (O. F. Muller)
PLATYHELMINTHES
Dendrocoelidae
Polycelis coronata Kenk
Procotyla sp.
ANNELIDA
Oligochaeta
Lumbriculidae
Tubificidae
Hirudinea
Glossiphoniidae
Helobdella stagnalis (Linnaeus )
Glossiphonia complanata (Linnaeus)
182
MOLLUSCA
Sphaeridae
Pisidium sp.
Literature Cited
Merritt, R. W.; Cummins, K. W. 1996. Aquatic insects of North America. Third edition.
Dubuque: Kendall Hunt Publishing. 862 p.
Pennak, R. W. 1989. Fresh-water invertebrates of the United States. Third edition. New
York: John Wiley & Sons. 628 p.
Stewart, K. W.; Stark, B. P. 1988. Nymphs of North American stonefly genera (Plecoptera).
Denton: University of North Texas Press. 460 p.
Wiggins, G. B. 1996. Second edition. Larvae of the North American caddisfly genera
(Trichoptera). Toronto: University of Toronto Press. 457 p.
183
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