Manual For Forest Tree Seed Orchard
Manual For Forest Tree Seed Orchard
Manual For Forest Tree Seed Orchard
of Canada
Gouvernement
du Canada
A manual for forest tree seed
orchard management in the
Canadian Service
Forestry canadien des Maritimes
Service forets
,cr
-
The program consists of three major elements research and development,
technical and information services , and forest resources development. Most
research and development work is undertaken in direct response to the needs of
forest management agencies , with the aim of improving the protection , growth, and
value of the region 's forest resource for a variety of consumptive and noncon-
sumptive uses: studies are often carried ouljointly with provincial governments and
industry . The Centre s technical and information services are designed to bong
research results to the attention of potential users, to demonstrate new and
improved forest management techniques, to assist management agencies in solving
-
day to - day problems, and to keep the public fully informed on the work of the
Maritimes Centre.
Cover photographs:
BOTTOM: Red spruce clonal seed orchard, Bowater Mersey Paper Company
A MANUAL FOR FOREST TREE SEED ORCHARD MANAGEMENT
IN THE MARITIMES
by
Government of Canada
1988
QAinister of Supply and Services, Canada 1988
The exclusion of certain manufactured products does not necessarily imply disapproval nor does the
mention of other products necessarily imply endorcement by the Canadian Forestry Service.
This publication reports research involving pesticides. Pesticides must be handled and applied properly.
All uses of pesticides must be registered by federal authorities and approved for use by provincial authorities
before they can be recommended.
UPDATES TO THIS MANUAL
Sections of this manual will be updated as required. If you want to automatically receive these updates,
complete the following and return to:
Scientific Editor
Canadian Forestry Service - Maritimes
Hugh John Flemming Forestry Centre
P.O. Box 4000
Fredericton, N. B.
E3B 5 P7
Please put my name on the mailing list to receive updates to “ A Manual for Forest Tree Seed Orchard
Management in the Maritimes”.
Name:
Affiliation:
Address:
I
ABSTRACT
Procedures are outlined for the establishment and management of seed orchards in the Maritime
provinces. Sections are included on site selection and orchard site planning, site preparation, orchard design,
layout and planting, control of competing vegetation , fertility management , protection, cone-crop enhance-
ment, and cone harvesting.
RESUME
Ce manuei indique les methodes a suivre pour bien etablir et amenager les vergers a graine dans les
provinces Maritimes. On y trouve des sections traitant de la selection et planification des sites des vergers,
preparation des sites, plans detailles de vergers et leur disposition sur !e terrain, plantation, suppression de la
vegetation concurrentielle, techniques d’amelioration de la fertility, protection , augmentation de la
production des cones, et recolte des cones.
ii
TABLE OF CONTENTS
PAGE
ABSTRACT
LIST OF TABLES vi
LIST OF FIGURES VII
ACKNOWLEDGEMENTS viii
PREFACE ix
CHAPTER 1 1-1
PLANNING 1-1
Site Selection , 1-1
General location . . 1- 1
Seedling seed orchards 1-1
Clonal seed orchards 1- 1
Specific location 1-1
Clonal and seedling seed orchards 1-1
Soils 1-1
Topography 1- 2
Insolation 1-3
Air drainage , 1-3
Accessibility 1-3
Isolation from foreign pollen . .. 1 -3
Past land use 1- 3
Orchard Site Plan 1-3
Area requirements 1-3
Seed production 1-3
Pollen dilution zone 1-5
Firebreak 1-5
Roads 1-5
Pond 1-5
Windbreaks 1-5
Equipment shed and offices 1-5
Layout 1-5
Suborchards , 1-5
Roads 1-7
Soil drainage 1-7
Windbreaks 1-7
Irrigation 1-7
Fencing 1- 7
Suggested Readings 1- 9
Literature Cited 1-9
CHAPTER 2 2-1
SITE PREPARATION 2 -1
Soil Drainage 2-1
Forested Sites 2-1
Seedling seed orchards 2-1
Clonal seed orchards 2-2
Field Sites 2- 2
Seedling and clonal seed orchards 2- 2
Cover Crops 2-2
Cover crop characteristics 2 -2
Fertility requirements for seeding . 2 -2
Seeding 2-3
Ill
PAGE
Maintaining fertility levels 2-3
Maintenance mowing 2-4
Literature Cited 2-4
CHAPTER 3 3-1
ESTABLISHMENT 3-1
Design and Spacing . . 3-1
Seedling seed orchards . 3-1
Design .. 3-1
Spacing 3-1
Clonal seed orchards ... 3-1
Design 3-1
Spacing 3-5
Layout, Planting and Tending 3-5
Seedling seed orchards . 3-5
Clonal seed orchards 3-5
Field Grafting 3-7
. .. .
Suggested Readings . .. . . 3-8
Literature Cited 3-8
CHAPTER 4 4-1
VEGETATION MANAGEMENT 4-1
Localized Control * * 4-1
General Control 4-1
Herbicides 4-1
Vision (glyphosate) 4- 2
Princep Nine-T (simazine) 4- 2
DCPA Dacthal 75W ( chlorthal dimethyl) 4 -2
Kerb SOW (pronamide) 4-2
Velpar L 4- 2
Combined sprays 4-2
Suggested Readings 4-3
Literature Cited 4-3
CHAPTER 5 5-1
FERTILITY MANAGEMENT 5-1
Assessing Site Fertility 5-1
Soil sampling 5-1
Assessing Soil Fertility 5-2
Soil acidity (pH) 5-3
Organic matter content .. .. 5-3
Cation exchange capacity .. 5- 4
Soil macronutrients 5- 4
Correcting Nutrient Deficiencies 5-5
Assessing Foliar Nutrient Levels 5-5
Optimizing Tree Growth 5-5
Suggested Readings 5-7
Literature Cited 5-8
iv
PAGE
CHAPTER 6 6-1
PEST MANAGEMENT 6-1
Assessing Cone and Seed Losses 6-1
Insects and diseases 6- 2
Monitoring Insects and Diseases 6- 2
Controlling Insects and Diseases .. 6 -2
Control of insects 6 -2
Control of diseases 6-3
Controlling Other Pests 6-3
Mammals and birds 6 -3
Vandalism . 6-6
6-6
Fire
Weather 6 -6
Suggested Readings . 6 -7
Literature Cited 6-7
CHAPTER 7 7-1
CONE CROP MANAGEMENT 7-1
Cone Crop Enhancement 7- 1
Fertilizers 7 -3
Type of fertilizer 7-3
Rate of fertilizer application ... 7-3
Timing of fertilizer application 7-3
Tree size 7 -3
Weather 7-3
Size of the current- year cone crop 7-3
Clone / family differences 7- 3
Cover crop 7-3
Root pruning 7-3
Girdling and strangulation .,. 7-4
Growth hormones 7-4
Topping trees 7-4
Pollen Contamination 7-4
Cone Harvesting 7-5
Assessing the cone crop 7-5
Time of collection * * *
»
•
7 -6
.
How to collect . . 7-7
Suggested Readings . 7-8
Literature Cited 7 -8
PAGE
APPENDIX VI ! MIXING YOUR OWN FERTILIZERS VII - 1
LIST OF TABLES
PAGE
Table 1-1. Estimates of seed orchard area requirements (adapted from Coles 1980b) 1- 4
Table 2-1. Soil fertility levels ( top 10 cm) for establishing a grass cover crop 2-3
Table 2-2. Date and rate of seeding for a seed orchard cover crop 2 -3
Table 5-1. Suggested soil fertility levels for seed orchards , Canadian Forestry Service
laboratory 5-3
Table 5-2. Suggested soil fertility levels for seed orchards, Nova Scotia Agricultural College
laboratory 5-4
Table 5-3 . Foliagesampling for nutrient analysis. The suggested numbers of shoots to be collected
is the minimum for the Canadian Forestry Service laboratory, and should provide ample
material for other laboratories 5-6
Table 5-4, Suggested foliar nutrient levels by species for seed orchards (From Mahendrappa
unpub! data ). Orchard trees can be considered as being adequately supplied with the
,
Table 5-5. Suggested schedule for fertilizing individual grafts in seed orchards 5-7
Table 7-1. Estimates of times of periods of initiation of potentially reproductive buds, and of
differentration of reproductive structures within buds, for species growing in average
conditions in the Maritime Provinces ( Powell 1983 with modifications by personnal
communications 1986) 7- 2
Table 7-2 . Evaluating cone crop seed yields from a cut -test 7 -6
VII
LIST OF FIGURES
PAGE
Figure 1- 1. Recommended areas forestablishment of clonal seed orchards in the Maritimes ( from
Coles 1980a ) 1- 2
Figure 3-3. Portion of a computer generated clonal seed orchard design ( COOL ) using 50 clones
and design type 5. 3-4
Figure 3 - 4. -
Layout patterns for clonal seed orchards: A - rectangular, B square , C-triangular. .. . 3 -4
Figure 5-1. A hypothetical seed orchard with its permanent road system used to subdivide the
orchard for soil sampling 5 -1
Major insect pests and theirfeeding times in Maritime tree seed orchards ( from Forest
Figure 6-1 .
Insect and Disease Survey, 1986., unpublished data ) , . 6- 4
Figure 7-1. The reproductive cycle of white spruce ( adapted from Table 7-1 ) 7 -1
Figure 7- 2. The reproductive cycle of jack pine ( adapted from Table 7 -1 ) . 7-1
Figure 7-3. Diagram of a pollen trap ( from Greenwood and Rucker 1985 ) . 7 -5
Figure 7 -4. Cone and seed maturation stages for eight Maritime tree species. Development stages
can be advanced 1 to 3 weeks with warm and dry weather and sites, and similarly
retarded on cold and wet sites or where growing seasons are naturally later e. g., the
Fundy shore ( from Smith 1985 ). 7-7
VIII
ACKNOWLEDGEMENTS
This publication would not have been possible without the assistance of many people. In
particular , we would like to thank Mr. Jim Coles for the initial impetus for a seed orchard management
manual .
We want to thank the following people for their valuable assistance and advice: Mr. R .D. Hallett -
.
Soils and herbicides; Dr . L .P. Magasi and Mr F. A , Titus - Insects and diseases: Dr. M.K . Mahendrappa
- Soils; Mr. T.R . Renault - Insects and diseases; Dr. G.R . Poweil - Tree phenology; and Mr. M.
Oudemans - artwork .
We would also like to thank the following people for reviewing the manuscript: Dr. D.P. Fowler,
Mr . T.J. Mullin, Ms . K .J. Tosh, and Mr. R . G. Wasser.
Finally, we wish to extend our sincere thanks to Nancy Hay and Patricia Gaboury for their
diligence and patience with us and the typesetter.
IX
PREFACE
The techniques used to plan, establish, and manage a seed orchard vary with the species involved
and the stage in the breeding program. In the Maritimes, for example, first generation seed orchards
for black spruce ( Picea ma /vanajMill ] B.S. P.) and jack pine ( Pinus banksiana Lamb,) , are seedling
.
orchards while second generation orchards will be clonal First and subsequent generation orchards
for most species will be clonal.
This manual outlines in detail , the steps required for planning, establishing, and managing tree
seed orchards in the Maritimes Region. The information is the best available, at present, but will
undoubtedly be updated as more knowledge is acquired.
1- 1
CHAPTER 1
PLANNING
Forest tree seed orchards are expensive to analyzed the climatic data for the Maritimes and
establish and manage and require continuous fund- identified the Annapolis Valley of Nova Scotia, the
ing. Therefore, careful planning of such a facility is southeast area of Prince Edward Island, and south-
necessary to ensure that funds are continuously western New Brunswick as having suitable climates
available. The planning process involves the selec
tion of a suitable orchard site and the development
- for establishing clonal seed orchards ( Fig. 1-1) ,
although there are also other suitable areas.
of detailed site plans incorporating soil, topography,
and area requirements for seed production, pollen Specific location
dilution , firebreaks, roads, and windbreaks .
Clonal and seedling seed orchards
Site Selection Soils. Texture strongly influences soil struc-
ture, erodibility and compactibility and because it is
Seed orchards are sound investments only if essentially unchangeable, it must be given high
they are fully productive. Environmental and edaphic priority when selecting an orchard site. A deep
variables of site influence seed production so care (30 to 50 cm) zone of sandy loam to loamy sand
must be taken to choose a site on which seed overlaying a friable subsoil presents the fewest
production will be maximized. A well chosen site problems with drainage and compaction. Heavy
can also minimize management problems. Twenty - clay soils must be avoided because compaction
five years of experience in British Columbia and the problems will result from the heavy and frequent
southeastern United States indicates that location is traffic in the orchard. Excessively drained sandy
crucially important in achieving abundant and soils should also be avoided because fertility levels
frequent cone crops. will be difficult to maintain ( low cation exchange
capacity) and trees will be prone to severe water
General location stress during dry weather.
Adequate soil drainage is necessary for on- site if water remains on the surface more than a day
accessibility in the spring and after heavy rains. It is following a heavy rain, artificial drainage will be
also important for the development of a deep and required.
extensive root system. Soil drainage should be
evaluated, and improvements ( e. g., drainage tile, Topography . The topography of the site
ditches) made before the orchard trees are planted influences insolation, air and water drainage, and
( APPENDIX l, Fig. I-2 ) . NOTE: Correcting a soil ease of equipment movement. Topography is also
water drainage problem 10 years after planting is essentially unchangeable, and is therefore an
virtually impossible without seriously damaging important consideration when selecting a seed
many of the orchard trees. orchard site. Select a site with a gentle (<5%),
uniform slope. A pit- and -mound topography is
The best time of year to determine if there are undesirable . Trees must not be planted in hollows or
drainage problems is in the early spring following depressions into which water drains. Planting in
snowmelt or after heavy rains . Ponding of water in ‘ wet ’ pockets will adversely affect tree survival,
depressions can be noted and the time required for growth, and cone production ( see APPENDIX I,
this water to soak into the soil should be monitored. Fig. I-3).
1-3
Insolation. In the northern hemisphere, south chemicals, if any, have been used and to have the
facing slopes receive the greatest amount of direct soil tested for potentially harmful residues. A poten-
insolation, consequently they are warmer. Simpson tial orchard site may have to be rejected because of
and Powell ( 1981 ) found that young black spruce the past use of arsenicals or high residues of
trees growing on south - facing sites produced more herbicides such as atrazine.
cones than those growing on sites with other aspects .
In the Maritimes, the only commercial labora -
Air drainage. Frost pockets must be avoided tory for testing for herbicide residues is the Atlantic
because conifer strobili are very susceptible to Provinces Pesticide Residue Laboratory in Kentville,
damage by late spring frosts . Do not locate a seed Nova Scotia (see Shreve and McCarthy 1985).
orchard at the base of a hill or in a depression where Testing for chemical residues is expensive
cold air will settle. Gently sloping sites with good air ( $120/ chemical, M. Shreve, pers. comm. Nov . 1986) .
drainage will be less likely to receive ‘killing’ spring
frosts than completely flat sites from which air does Orchard Site Plan
not drain readily.
Having selected a suitable site, a detailed plan
Accessibility . The orchard must be accessible encompassing all the long -range goals is necessary
and operable during ail seasons of the year par- for a successful and productive orchard complex .
ticularly in early spring. In seedling seed orchards, The components of planning an orchard are dis-
stumps can pose a problem for early mechanized cussed below. Although most items apply specif-
cone collection. A well kept road system MUST be ically to clonal seed orchards they also can be
maintained. It is advantageous to locate an orchard applied to seedling seed orchards. APPENDIX I , Fig.
near a nursery or field office. A close source of I-4 presents a general site plan, which should be
manpower and equipment facilitates monitoring the made BEFORE orchard establishment begins.
orchard and helps control access.
Area requirements
Isolation from foreign pollen. An orchard
should be located on a site where contamination by There are many considerations in determining
pollen of the same or closely related species is total area requirements, such as the number of
minimal . Locating seed orchards either outside the orchards (if more than one) to be combined at one
natural range of the species or in a warmer climate complex, and allowances for roads, drainage
where the orchard trees are out of phase with the ditches, and ponds, equipment storage or main-
local trees have been suggested as potential solu- tenance areas, poor or unusable land , pollen dilu-
tions to the pollen contamination problem ( Gansel tion zone, firebreaks, and windbreaks. These could
1973; Sarvas 1970; Werner 1975). In some areas of double the total area required for an orchard com-
the Maritimes, it may be impossible to locate an plex and must be considered during the planning
orchard outside the natural range of a species such and site selection stages.
as white spruce ( Picea glauca [ Moench ] Voss).
Therefore, a compromise should be adopted such Seed production
as locating the orchard as far as practicable from There is at least a 7 to 10 year lag between
natural stands of the same species, removing trees initiation of a tree improvement program and the
of the same species from the surrounding areas, and production of quantities of genetically improved
locating orchards on warm sites. seed . Consequently, long -range planning of a
reforestation program is a must . The area of seed
Past land use. The history of a site being orchard required must reflect the planting program
considered for an orchard should be known. For 20 to 30 years hence.
land previously farmed, it is vital to determine what
1- 4
The area of seed orchard necessary to produce Table 1 -1 summarizes the actual area of orchard
sufficient quantities of seed for a particular species to be planted to produce one million plantable seed -
can be calculated. The number of seedlings required lings a year by species and is an updated version of a
annually and the quantity of seed produced from a table found in Coles ( 1980b } . APPENDIX II explains
unit area of orchard need to be determined. There the assumptions and calculations. Two important
are, however, a number of other factors involved in points should be emphasized 1) cone production
estimating area requirements. per tree is an estimate falling between early produc -
tion and late production ( time orchard is phased
1. Seedling production system ( bare - root vs.
out) and 2) trees per unit area is the number after
container ) .
final rogueing. Hence, for clonal seed orchards, in
2. Periodicity of cone crops. particular, before rogueing, seed production will
3. Number of trees or grafts per unit area. probably exceed those estimates presented here.
4. Number of cones produced per tree or graft.
5. Number of viable seed per cone.
Table 1-1. Estimates of seed orchard area requirements (adapted from Coles 1980b)
Method of Area required for
Species Interval seedling 1 million plantable
Type of seed Number of Sound Trees / ha between cone production seedlings /year
orchard (S. O. ) cones/ trees1 seed/cone ( spacing) 2 crops ( S.S . - sound seed) (ha )
1The number of cones per tree will increase with age and size. The figure given is an estimate which falls
between early production and that of maturity of the orchard.
2The number of trees per hectare will decrease from establishment to maturity. The figure given represents
the number remaining following final rogueing.
1-5
Roads Layout
Easy access throughout the orchard is neces -
sary. Major roads will normally separate the sub-
Once the areas required for the various com-
ponents have been calculated, a map of the site
orchards in a complex. Secondary roads will sepa-
incorporating the appropriate components (Fig. 1-
rate blocks within the suborchards and occur around
the outer perimeter. A strip 10 m wide for major 2) should be drawn. The map should be drawn to
scale and suitable to overlay other maps containing
roads and 6 m for secondary roads ( sufficient to
accommodate equipment turning ) should be soil and topographic information, and drainage
plans ( see APPENDIX I, Figs. 1-1 to I-3) .
allowed.
Suborchards
Pond
If an orchard complex containing several spec-
An orchard complex cannot be operated suc-
ies is being considered, the positioning of the
cessfully without an adequate supply of water for
individual species in the suborchards should be
irrigation, fire protection, chemical spraying, etc .
considered. Each suborchard must be of sufficient
The Atlantic Committee on Agricultural Engineering
size to produce enough pollen and to accommodate
publication titled "Farm Ponds” (Higgins 1984)
proposed expansion . The minimum size of a sub-
provides all the necessary information for pond
orchard should be 3 to 4 ha and be arranged so that
construction and maintenance. About 0.5 ha should
the prevailing wind at the time of pollination blows
be allowed for a pond area near the centre of the
parallel to the long axis thus promoting within-
orchard complex. An agriculture engineer from a
orchard pollination. Species that readily hybridize
provincial Department of Agriculture can assist in
( e.g., black spruce and red spruce ( Picea rubens
the planning and designing of a pond. Ponds con -
Sarg.) should not be planted in the same complex.
structed in the Maritime Provinces must also meet
any specifications and regulations laid down by the
appropriate provincial Department of Environment.
A well or river is an alternate source of water.
cr>
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- ^r .*v
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- 1995 \ I
1997 SUB ORCHARDS
mmmmmm
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BLOCK AREA ( HA )
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m
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WP - 2
•• * > *' >
t
4
\ /
i
1
5.0
i
t TOTAL 10
/ !
;
i
»*
\
i k
i WS - 1 3.0
‘ \
*1 W?i.*
i h t
WS - 2 3.9
- ^A
>
WS - 4 * <)*?$?*?-
i < k !
l WS - 3 I k
k j * *.•
WS - 3 2.5
»
—
\
WS - 4
i
-/ / t 2.6
' J• -iat
:k
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White \
5 k * k '
:
TOTAL 120
< /* White Wi®f&
t
k
t;
Black Spruce 10.5
K*
> Spruce \
Jack Pine 8.5
field k
Pine
*/ \ \
Pond Area 1.2 '
WS - 2 «
WS - I WP - I WP - 2
i feifsi
!
i
k
i
i
t
TOTAL 42.2
KS@S
i l!
*/ -i. \ .»! \
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t/ m k ii
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i \
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i
i
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k
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i
ilGHV I i
^ 1000
V.
field
Concentration
of cations ( ppm)
K Beneficial to plant nutrition .
Ca * 120 Higher levels prevent P injection and increase pre -
cipitation in irrigation lines ( see fe )
Concentration of
trace elements ( ppm )
Fe 50 May cause precipitation of phosphates and clog
nozzles
Mn 0.2
Zn 20
R 05 *.2.0 >2.0
Mo 0.01
Concentration of
anions ( ppm )
SO , :480
Calculated indices
SAR < 10 18 18
*
ASAR * .3 :- 3 indication of potential Na toxicity
Gansel, C. R. 1973. Should slash pine seed orchards Werner , M. 1975. Location, establishment and man-
be moved south for early flowering? Pages 310- agement of seed orchards. Pages 49-57 In Seed
.
316 In Proc. 12th South . For Tree Improv . Conf ., Orchards. Faulkner R . (ed.) For. Comm. Bull. No.
Baton Rouge, LA, June 12-13, 1973. 54.
CHAPTER 2
SITE PREPARATION
Proper site preparation on ail orchard sites titled “Farm Drainage in the Atlantic Provinces”
facilitates subsequent management operations ( Gartley et al. 1986) provides valuable information
such as layout of planting spots, planting, fertilizer on soil drainage systems. Contact a Department of
and chemical applications, and cone collection. Site Agriculture drainage engineer for advice on the
preparation differs forseedling and clonal orchards appropriate method ( s ) to solve soil drainage
both in methods and intensity. APPENDIX III problems.
includes a table for summarizing and recording the
various operations. Forested Sites
If a hardpan is found during initial site assess- Site preparation for seedling seed orchards
ment, the type of problem, if any, that it creates must should not change the environment appreciably
be determined. A hardpan at a depth greater than from that of a normal planting site because the trees
1 m will do little to limit rooting directly but may are evaluated for differences between families
affect it indirectly through impeding water drainage. ( when rogueing ) and for the production of seed.
Hardpans at depths greater than 1 m will, for all Seedling seed orchards are best established on
practical purposes, be too deep to be broken up. cutover forest land. Preparation should aim at
The options for correcting water drainage problems improving the uniformity and operability of the site.
are as follows. A major objective of any type of scarification is to
1. Avoid them. mix the organic matter layer with the mineral soil
Ensure that there is sufficient extra area to meet and not expose mineral soil or strip off the organic
requirements and that these abnormal areas do matter layer. Slash should not be piled and burned
not affect the remaining area, e.g., wet areas on the site, except where roads are planned,
may break up the orchard creating problems in because burning alters soil nutrition regimes.
operation.
During the cutting operation, stumps should be
2. Rectify them,
cut as low as possible. The slash can be removed
a. By deep subsoiling ( break up hardpans).
from the site using a brush rake such as an Eden
b. By installing ditches or drainage tile ( remove rake or Raumfix rake . An experienced equipment
excess water ) .
operator is required because the soil should be ieft
c. A combination of a and b .
intact. If slash is not removed then double crushing
or a double pass with Drum choppers or a Rome
Subsoiling to a depth of 50-100 cm, using a
Disc wilt break up and flatten it. The Madge Roto-
large bulldozer equipped with a ripping claw, breaks
clear , which acts like a rototiller, chews stumps and
up the hard pan. A grid pattern at 3-4 m spacing will
siash and mixes them with the mineral soil to create
provide good coverage of an area. Drainage tile may
be necessary if there are large depressions on the
-
a garden like site. Stumps must be no higher than
12 cm above the ground and be at least 6-12 months
site. A subsurface drainage plow using a laser grade
old.
control system can be used to install plastic drain-
age tubing. A large blade, with a hollow chute into
Competing vegetation must be controlled
which the tubing is fitted, is pulled through the soil.
As the plow moves, the tubing is drawn through the
before it affects the growth of the orchard seedlings.
The most effective and cost efficient time to begin
chute into the soil at the bottom of the blade. The
weed control, is BEFORE planting . However,
laser establishes a pre-determined sloping refer-
because unwanted vegetation does not usually
ence plane over the site allowing drains to be grow until after harvesting, such treatments may not
installed with great accuracy . The Atlantic be applied unfit the seedlings have been planted
Committeeon Agricultural Engineering publication (see CHAPTER 4 for herbicide treatment).
2-2
Clonal seed orchards vegetation growing on the field, the entire area
should receive an application of herbicide in late
The sole purpose of clonal orchards is to spring or early summer ( see CHAPTER 4 for herbi-
produce abundant quantities of seed. Therefore cide application) . About one week following the
everything possible should be done to favor early herbicide application the site may be plowed and
and abundant cone production . harrowed. Fertilizer and lime amendments, as recom-
mended from results of a soil analysis, should be
Slash and stumps should be removed from the applied and harrowed into the soil. This operation
.
site using a root rake As much soil as possible may not be necessary if the site is to be used for a
should be shaken from the roots before they are seedling orchard unless certain nutrients are
removed. Slash and stumps should not be burned limited. If during the summer fallow period, weeds
on the site, except where roads are planned, and grass reestablish on the site, a second herbicide
because of the effect of burning on soil nutrition. application followed by harrowing may be neces-
Large boulders can also be removed from the site at sary, This should be done at least one to two weeks
this time. before the cover crop is sown. Frequent harrowing
during the summer months may be sufficient to
The site should now be harrowed to smooth the control regrowth of weeds and grasses.
surface and raked to collect surface rocks and
pieces of roots and slash in windrows where they Cover Crops
can be removed with a rock picker. Following this
operation, large depressions can be filled and An essential part of a clonal seed orchard is a
mounds removed to produce a flat surface. Plowing permanent grass cover crop. Clare et at. (1984)
the soil to a depth of 25 to 30 cm will mix in the provide guidelines for the establishment and man-
organic matter layer and when harrowed will further agement of the permanent cover crop.
loosen the soil and make it more manageable. Rock
picking may again be necessary. Cover crop characteristics
Fertilizer and/ or lime amendments should be A seed orchard cover crop should be long lived,
applied at this time, based on the results of soil and produce a thick sod that is able to prevent
sample analyses ( see CHAPTER 5, Assessing Site erosion and withstand traffic. If should also exhibit
Fertility ). Harrowing will help to incorporate the short growth (infrequent mowing), low fire hazard
amendments into the soil and prepare the site for ( thatch remains green) , contain no clover (rodent
sowing the cover crop. control ), and be aesthetically pleasing.
Seedling and clonal seed orchards Growing and maintaining a grass sod requires
regular fertilizing with N, P, and K . Some soils
When a field site is selected for a seed orchard, require additional Ca and Mg as determined by soil
the site preparation procedures will be similar for analyses. Fertility levels of the orchard site MUST
both types of orchards, but less intensive for seed- be determined BEFORE seeding the cover crop
ling orchards. Seedling orchards are not usually ( Table 2-1).
established on fields because such locations are not
typical planting sites for reforestation programs. Lime is usually required in the year of estab-
However, occasionally field sites are used because lishment. Lime and fertilizer should be mixed into
preparation costs may be lower and initial manage- the soil to a depth of 8-12 cm before seeding. No
ment may be more economical. benefit comes from deep incorporation as most
grass roots are shallow.
Before operations begin, the site should be
assessed to determine if land forming is necessary. Fertilizer and lime can be spread by broadcast
If required, it should be completed before plowing or band type ( Gandy ) spreaders. The band type is
and harrowing. If a hardpan, exists, it must be recommended as the fertilizer can be applied more
broken up by methods discussed earlier. If there is accurately to the grass and not on the trees.
2-3
Table 2-1. Soil fertility levels ( top 10 cm) for establishing a grass cover crop
Organic
matter P205 K 20 Ca Mg
pH ( %) ( kg/ha) ( kg/ha) ( kg/ha) (kg/ ha)
1 At or below minimum values, grass will be less vigorous and may die in stressed areas.
Rate
Time Fertilizer (kg/ha)
Years 2 -5
Spring 34-0-0 -
50 70
Summer 1 34-0-0 50-70
After year 5, sod in poor shape, years of heavy traffic , or low P & K
' Summer means late June to early July, not after mid-July. Always do a soil test and use this table only as a
guide.
Maintenance mowing
Literature Cited
CHAPTER 3
ESTABLISHMENT
This phase of seed orchard development in- magnitude allow for reasonable selection intensi-
volves selecting a design , spacing, and layout that ties during rogueing and the production of out-
are easy to implement, best satisfy genetic con - crossed seed. When laying out the orchard, a 5 -m-
siderations for seed production, and provide ease of wide strip should be left unplanted at every 15 th row
orchard tending. Extreme care and attention to for access roads. Stakes are placed at the beginning
detail are ‘musts' during the planting and mapping and end of each row and at every 50th planting
stages to ensure good survival and to prevent errors. position within rows. These subdivide the rows into
Tending and maintenance following planting are subsets providing tie- in points ( Fig. 3-1A ).
important to promote quick establishment and early ,
rapid tree growth. APPENDIX IV , Table IV-1 is a When using the block design, blocks must be
form for recording various operations on an annual sufficiently large (50 X 50 m) because area is
basis. required for access roads. Seven to nine seedlings
per family should be planted in each block depend-
Design and Spacing ing on spacing . A minimum of 150 families is
necessary to ensure a sufficient number of unre-
The major objective of a seed orchard design is lated trees remain after final rogueing . To ensure
to maximize out -crossing and panmixis ( random that ample area is planted at any one time, 12-15
pollination) while minimizing self -pollination and blocks are recommended (3-4 ha) . A 5-m- wide
related mating. It is also important to provide for access road is left around each block but no
future expansion and rogueing. Spacing of the provision is made for vehicle access within the
orchard trees should take into consideration blocks. Both endsof the rows are staked ( Fig. 3-1B) .
planned rogueing intensity and easy movement of
equipment. Spacing
Spacing must be close enough to allow for
Seedling seed orchards planting many seedlings per unit area but not so
close that competition for growing space forces too
Design early and intensive rogueing. Conversely, if too
A design incorporating planting a large number wide a spacing is used, the orchard cannot be
of families with many seedlings of each family at rogued as heavily as may be required to maximize
close spacing is recommended for seedling seed genetic gain or too few trees may be left , thus
orchards. The close spacing allows for two to three reducing seed production per unit area .
rogueings with a final rogueing intensity of 85 to
90%. The intensity of each rogueing depends on the The 2 x 1-m spacing generally used in the
need to maintain a full, open crown on the remain- Maritimes, appears to be adequate for black spruce,
ing trees, maximize genetic gain , and assure easy but for jack pine which grows faster and produces a
movement of equipment. larger crown, 1.25 x 2.50 m may be more
appropriate.
The general design that is being used for
seedling orchards in the Maritimes is thesingle-tree Clonal seed orchards
plot. In this design one or more seedlings from each
family are randomly planted in each row or block of Design
the orchard. For clonal orchards, a more sophisticated
design is necessary because fewer unrelated indi-
Using the row design, the number of rows is viduals are planted and rogueing is at a lower
determined by the average number of seedlings intensity than in seedling seed orchards. Forty to 50
available per family. A minimum of 100-150 families clones are sufficient for a clonal orchard or in
with 100 seedlings per family should be planted at separate blocks if the site is large ( greater than 4
any one time (minimum area of 3-4 ha, if no ha). This number will be reduced by 50% by rogue-
additional area is to be planted) . Numbers of this ing and miscellaneous losses.
3- 2
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X X X X X
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• = stake * = seedling
Clonal orchards will be rogued primarily on the Using the random orchard design, each ramet
basis of results from control pollinated progeny of each clone has an equal opportunity to be
tests. Clones that do not produce large numbers of selected for any given planting position. The one
cones will be culled. major restriction imposed is that at least two dif -
ferent ramets must separate those of the same
Many designs have been proposed for clonal clone. This design may be implemented by com -
seed orchards. The ramets can be planted either pletely randomizing all the available ramets of all
systematically or randomly. When an orchard lay- clones among all available planting positions, or by
out is designed systematically the ramets are placed dividing the area into blocks each sufficient in size
in a predetermined pattern (Fig. 3-2). Although such to contain one ramet of each clone or a multiple so
designs maximize outcrossing, panmixis is much there are an equal number of ramets per clone.
reduced and the spacing after rogueing may be
unacceptable.
3-3
1 2 9 1 2
4 5 12 3 5
*• >
7 8 4 7 6
10 11 12 8 10 11
Block 1 Block 2
1 2 3 4 5 6 7 8 9
4 5 6 7 8 9 1 2 3
7 8 9 1 2 3 4 5 6
1 2 3 4 5 6 7 8 9
A computer program called COOL (computer Figure 3-3 illustrates a portion of an orchard
organized orchard layouts) , developed by Bell and designed by this program . The diagram shows that
Fletcher ( 1978) to produce random layouts, is being when a design type of 5 is chosen, a ramet of clone
used in the Maritimes by the Canadian Forestry 50 is separated from all other ramets of that same
Service - Maritimes, Fredericton; P.E.I. Department clone by at least four rings of different clones. A
of Energy and Forestry, Charlottetown; and N.S. design type of 5 should be considered as a minimum
Department of Lands and Forests, Debert . The when this program is operated to maximize dis-
design is based on the permutated neighborhood tances between ramets of the same clone .
concept and randomized with two restrictions:
proximity of two ramets of the same clone and Three layout patterns may be used: rectangu-
repetition of the direction of two adjacent clones. To lar, square, or triangular ( Fig. 3-4). The triangular
run the program , the following information is pattern allows more efficient use of growing space
necessary. for the tree crowns than does the square. The rows
in the rectangular design should be oriented east-
1. The dimensions of the orchard as defined by a west to allow the crowns maximum light exposure
specified number of rows and columns. on the south side.
2. The number of ramets per clone.
3. Design type that specifies the number of
planting positions in any direction by which a
clone is isolated and within which no other
ramet of the same clone can appear.
4 . The number of times two clones can occur in the
same immediate position relative to each other.
3- 4
21 44 22 11 43 25 35 19 45 6 46 2
34 15 25 1 5G 37 45 29 18 36 21 42 12 37 34
31 33 16 49 3 4 14 23 28 8 33 49 44 25 35
45 48 19 10 17 31 20 46 11 38 1 26 39
12 28 27 24 8 42 / 30 7 47 16 40 \ 30 18 4 23
*
* .
38 5 47 43 / 22 S 21 29 /44 49 15 \ 6 3\ 5 32 28
«
1
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31 44 16 41 23 i
1
12 ;'45
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24 36 14 17 26 ; 31 ! 5 '38 5 o [_ ]
26! 8 ! 10 ! 21 37 35
*
_ /22
%
* *
2?\J6 41 7
45 15 27 4 34 I 19 i30
7 20
38 32 3 37 24 \ 42 *» 49 17 3 33 2,'' 14 /25 43 23
10 1 46 6 36 26 \ 11 47 31 23 34 / 5 44 30 6
35 8 12 45 43 28 5 12 18 10 29 24 35 37 27
24 15
® 30 22 41 48 Ho ] 1 4 36 21 3 33 41
Figure 3-3. Portion of a computer generated clonai seed orchard design ( COOL ) using 50 clones and design
type 5.
X X X X X X X X X X X X X
X X X X X X X X X X X X X
X X X X X X X X X X X X X
X X X X X X X X X X X X X
A B c
Figure 3- 4. Layout patterns for clonal seed orchards: A - rectangular, B - square, C - triangular.
-
35
design map, a permanent metal tag containing the The size and fragility of grafts dictate careful
clone number should be attached to each stake. handling - KEEPTHEM IN AN UPRIGHT POSITION.
Other information, such as row number, position Large plastic or metal containers labelled with the
within row, block number, year grafted, etc., may row and planting location can be used. The roots
also be inscribed on the tag. Large wooden stakes must be protected and kept moist by covering them
reduce the aesthetics of such orchards, especially with wet peat moss. Grafts may also be individually
when the grafts are small. Short metal stakes or placed in buckets or plastic bags.
heavy guage wire to which the tags are attached can
be used. A large auger or posthole digger mounted on a
farm tractor is a good combination for digging holes
Herbicide can be applied to an area, up to 1 X 1 but care must be taken to reduce compaction of the
m, around each planting position or a strip down the sides of the holes. If this happens, the sides should
rows before planting the grafts. If the herbicide is be loosened by making vertical cuts with a shovel
applied after the grafts have been planted and they before the grafts are planted. NOTE: Do not bore
are actively growing, extreme care must be taken to holes when the soil is wet.
prevent the herbicide from contacting the trees (see
CHAPTER 4 for suitable herbicides) . Carefui outplanting minimizes losses and plant-
ing shock . The roots must be spread out in an
Only vigorous grafts with heaithy shoots and adequate sized hole and planted at the same depth
buds should be planted in the orchard. Grafts from as they were previously growing in the nursery or
winter greenhouse grafting may either be trans- pot. Forgraftsin pots with roots fully occupying the
planted to the nursery in the late spring or early potting medium, make several vertical cuts through
summer or retained in pots until they are large the roots with a sharp knife to help promote lateral
enough to be planted in the seed orchard. Current- root growth.
year grafts that have grown well can be transplanted
to the seed orchard the same year. Square pots, It is often necessary to have the grafts staked to
4.5 L, with root-trainer ridges on the inside walls are prevent snow and ice damage during the first two or
suitable for growing grafted material for 1 to 3 years three winters in the orchard. This also encourages
with a minimum of root deformation. Such material some grafts to grow upward to form a tree rather
should be overwintered in an unheated, shaded, than to grow in a branch-like fashion (topophysis).
sealed greenhouse to avoid damage. Potted grafts All grafts not having a topophytic growth habit
are easier to transport to the seed orchard for should be untied during the summer months. This
planting than nursery transplants, and may be encourages the development of a more sturdy tree.
planted one year sooner because planting shock is
minimal and root loss and deformation are elimi- Grafts must be carefully pruned for two or three
nated or reduced. The current trend in the Maritimes years to remove all the branches from the root stock,
is away from transplanting grafts to the nursery bed leaving only the grafted scion. Pruning should be
and subsequently to the seed orchard, to planting initiated at the greenhouse/nursery but completed
current-year grafts into the orchard and overwin- in the orchard. Pruning is best done in the spring
tering the remaining grafts for orchard planting the before the buds have flushed and can be repeated in
following spring. early August. Large branches should be removed
first. If large branches are left for several years, the
Grafts should be lifted from the nursery before size of the wound following pruning increases
the buds flush. This must be coordinated with the stress on the graft. If all large branches cannot be
phenology of the species. Tamarack , for example, removed at one time, those remaining may be
should be dug as soon as the ground thaws, whereas pruned back . When removing whorl branches,
red or black spruce can be ieft in the ground longer. never remove more than two at one time and remove
Before lifting, each graft should be tagged with ones opposite each other. Removing most or ail the
plastic nursery tags or permanent metal tags record- whorl branches at one time creates too much wound
ing the clone number. When lifting, as much soil as area thus promoting tissue desiccation and even-
practical should be left on the rootball. The roots tually girdling. Branches should be pruned flush to
and shoots can be pruned as required for handling. the branch collar ( Fig . 3-5) . Damaging this collar by
Figure 3- 5. Proper pruning method on conifer grafts .
pruning too close retards the tree’s natural wound DO NOT apply granular fertilizers until a month
healing processes and allows the entry of micro- after transplanting the grafts in the orchard. Fer-
organisms ( Shigo 1985 ) . Treatment of pruning tilizer is not usually required the first season fol-
wounds is not necessary. Grafts growing vigorously lowing planting as there should be ample nutrition
can be pruned more heavily than slow growing in the potting medium or soil within which the roots
ones. Keep the scion dominant at all times. NOTE: havegrown. However, if fertilizer is to beapplied the
Use discretion when performing this task . same year as transplanting, it should be placed
around the outer edge of the planting hole before
Following planting, a mulch layer of peat moss the mulch is spread . Ammonium nitrate and 10-10-
or sawdust around each graft increases survival and 10 in a 2:1 mix by volume at the rate of 30 g per graft
early growth. Mulch provides a layer of insulation is sufficient and will not harm the roots if applied
moderating soil temperatures at ground level and carefully.
helps retain soil moisture and nutrients while reduc-
ing competing vegetation. Well decomposed saw- Field Grafting
dust should be used as it does not immobilize soil
nitrogen to the same extent as fresh sawdust. Field grafting may produce a healthy, fast
Ground-up cones should be avoided because there growing tree and result in early cone production as
inevitably will be some viable seed present and the it eliminates transplanting the grafts to the orchard.
cones will also immobilize nitrogen from the soil as It also extends the grafting season. The major dis-
they decompose. The mulch should be applied in a advantages are that grafting must be done outdoors
40- to 60-cm-diametercircle 5-cm-thick , around the often under difficult conditions, success is usually
grafts but should not be in contact with the stem. lower than with greenhouse grafting, and growth is
Too thick a layer may encourage nesting or bur - poorer the first two to three years.
rowing by mice or other rodents or attract insects.
3-8
This technique, although not used extensively Tidswelt, K ; Dufour, A . 1984. Field grafting of white
in the past, shows promise for use in the Maritimes. spruce in the Maritimes. Marit. For. Res. Cent.,
Complete sections /blocks of an orchard should be Tech. Note No. 119.
established atonetime. Combining field andgreen-
house grafts often complicates early tending and Literature Cited
management . The root stock, two seedlings per
planting position, is established one to two years Bell, G . D.; Fletcher , A.M. 1978. Computerorganized
prior to grafting. The grafting may be conducted o r c h a r d l a y o u t s ( C O O L) b a s e d o n t h e
during mid - to late spring ( May). NOTE: Be careful permutated neighbourhood design . Silvae
not to leave any ungrafted rootstock. Genet. 27:223- 225.
For spring grafting , use dormant scions col- Hallett , R . D.; Smith, R .F.; Burns,T .W. 1981. Manual
lected in March and stored frozen in snow. The for greenhouse grafting of conifers in the Mari-
scions are grafted using the side-veneer grafting times. Dep. Environ,, Can . For. Serv., Marit. For.
technique ( see Hallett et al. 1981) , Protection from Res. Cent., Inf Rep. M- X -117.
,
CHAPTER 4
VEGETATION MANAGEMENT
There are numerous types of herbicide appli- DCPA Dacthal 75 W (chlorthal dimethyl)
cators, which can be used to apply herbicide around
trees, minimizing the risk of herbicide accidentally Dacthal is effective in controlling annual
contacting foliage . Sprayers equipped with grasses and some broadleaf weeds but is not as
"shrouds ” can be used when the seedlings are effective forperenniai weeds or grasses ( Kersting et
young to protect them against the spray. For larger al. 1983). It is a mainstay in many nurseries for
trees, a field sprayer can be modified with a shroud control of weeds in seedbeds. Like other preemer-
to direct the spray to the ground and under the lower gence herbicides it must be applied early in spring
branches. Wick-type applicators can also be safely before weed seed germination or growth commen-
used but are time consuming. Descriptions of the ces. A major advantage of DCPA is that there are
equipment available and their calibration can be generally no phytotoxicity problems with most of
found a handbook published by the B.C. Min. our native conifers (Hallett and Burns 1984).
Environ, (1986) ,
Kerb 50 W (pronamide)
Several of the chemicals that potentially could
be useful for vegetation control in seed orchards are If the buildup of simazine residues or its toxicity
briefly described below. is of concern then Kerb is an alternative. Kerb,
applied to the soil as a spray in the fall, controls
Vision (glyphosate ) many overwintering annuals and perennials. It is
applied only on cold soil because at warm soil
Vision (formerly called Roundup) was regis- temperatures, it readily volatilizes. By summer of
tered in 1984 for forestry use. It is effective against the year following application, most of the Kerb will
many species of weeds and hardwoods, is of low have dissipated. Kerb is absorbed through the roots,
toxicity to wildlife, fish, and humans, and is quickly thus to be effective, moisture from rain, irrigation, or
inactivated and degraded in soil. Woody species are snowmelt is essential to move it into the rooting
most sensitive in August or September before frost zone.
( Kersting et al. 1983). During the growing season,
Roundup can be used as a knockdown herbicide, Velpar L
but it must be applied around the tree seedlings as a
directed spray and not applied on the crop trees. Velpar L was registered in 1984 for woodland
use. It is effective against grasses, broadleafed
Prlncep Nine- T (simazine) weeds, and woody perennials. At low application
rates, black and white spruces and jack pine may be
Simazine inhibits seed germination. Therefore, planted immediately after application, but ALL
it must be applied either when the ground is free of OTHER conifer species should not be planted until
weeds or in combination with another herbicide the following year regardless of the application rate
such as Vision which provides the knockdown. used (Teskey and Boyer 1984).
Simazine is best applied when trees are dormant as
it has considerable soil residue activity. Larch and Combined sprays
container stock are especially sensitive to damage
by simazine while spruces are more sensitive than Some herbicides can be combined to achieve a
pines and fir. Conservative rates should be used for broad spectrum of control for a long period. For
the more sensitive tree species and stock types. At example, Vision and Princep can be combined, the
low application rates, grass control may not be former being used as a knockdown spray while the
complete. Because of its residual activity, annual latter provides residual weed control. Princep can
applications may not be necessary. Soils should be be combined with Dacthal to provide a broad
tested for toxic buildups of simazine if repeated spectrum control of grasses and broadleaved weeds
applications have been used (see CHAPTER 1, Past throughout the growing season (Hallett and Burns
land use) . 1983) .
4-3
Weed Science Society of America. ( W.S.S. A .) . 1983. Sims, H.P.; Mueller-Dombois, D. 1968. Effect of
Herbicide Handbook. 5th ed., Weed Sci. Soc. grass competition and depth to water table on
Amer., Champaign, III., 515 p. heightgrowth on coniferous tree seedlings. Ecol.
49:597-603.
Literature Cited
Teskey , K.K .; R. Boyer, L. G. 1984. Velpar L is now
British Columbia Ministry of Environment. 1986. registered for woodland areas. Dupont Canada,
Handbook for pesticide applicators and pesticide Industry News.
dispensers. A.V. Miller and S.M. Craig [ Eds.] .
Pesticide Control Branch, B.C. Min. Environ .
5 -1
CHAPTER 5
FERTILITY MANAGEMENT
Soil fertility will affect the quantity and quality It is not practicable to try to manage a seed
of seed produced in the orchard. The control of orchard on an individual-tree basis, nor should a
orchard fertility can be regarded as a two-stage 100 ha orchard be managed as one entity. There-
process: assessment and maintenance. The assess - fore, workable management blocks, that can be
ment stage involves determining the fertility status treated as homogeneous units must be identified.
of the site and alleviating any deficiencies The . To do this, blocks/ areas are identified which can be
management stage involves maintaining nutrients managed separately. The management blocks are
at optimal levels for both tree growth and cone usually determined by some logical subdivision of
production . Annual fertility management opera- the orchard such as species, suborchards, or year of
tions should be recorded ( APPENDIX IV ) . establishment within a given species ( Fig. 5-1).
Assessing Site Fertility In the hypothetical seed orchard ( Fig. 5-1) , four
management units are delineated by roads. Infor-
Soil sampling mation on the type ( s) of soil is obtained from soil
pits ( P ) and fertility samples ( * ). Before establishing
Soils in an orchard must be systematically the orchard, this information is used to identify
.
sampled Soil pits should be dug and bulk soil differences in soil types both between and within
samples collected for nutrient assessment. the management units.
—— ©- * * © * *
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B 3
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C« 12.0 ha ©* * © * * © * * © * * * *© *
D 3
I0.0 ha
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;
*© * *- * * -* * ®* **
® Soil pit
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* IPood} * * * * * * * *
'
fertility © ©
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/ *
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* © * * © -* * ©* * © * - * ® * * ®- * * I©*1
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for soil
Figure 5- 1. A hypothetical seed orchard with its permanent road system used to subdivide the orchard
sampling.
5- 2
Soil pits are systematically located to represent 2. Using a soil auger , take cores to a depth of 15 to
soils from the entire orchard area. The number of 20 cm. Small diameter augers, e.g., 2 cm, or
soil pits required increases with increasing variation larger sizes can be used. THE SAME NUMBER
on the site. Soil types often change with slope OF SAMPLES SHOULD BE COLLECTED
position, distance from a body of water ( old river- REGARDLESS OF THE SIZE OF THE AUGER .
bank terraces), etc. The three pits located around 3. Mix all the samples from each sampling area,
the pond in Fig. 5-1 are necessary to determine how removing large roots, clumps of sod, etc.. Use a
far the influence of the low wet area extends. The CLEAN “ ship n’shore” container in which to mix
boundaries of any other abnormal areas should be the soil. DO NOT use old oil barrels or similar
similarly delineated, e.g., hardpans. NOTE: This containers that might contain residues which
assessment is done BEFORE orchard trees are could contaminate the soil .
planted ( see CHAPTER 2, Soil drainage). 4. If a large diameter (10 cm) sampling auger is
used the total amount of soil collected will be
Sampling to determine variation in a single much more than is needed for analysis. Mix the
orchard block should be done in one year. DO NOT soil thoroughly and takea 1-Lsubsample random-
collect a small number of samples and if the varia- ly from the total soil sample.
tion is high, collect more samples the following 5. Place each sample separately in a heavy paper
year. If samples are collected in year 1, and subse- .
bag (sugar bags are ideal) Label the bags with
quently the site is disturbed , e.g., plowed and waterproof ink , and map the exact locations in
harrowed, nutrient levels of soil samples collected .
the orchard where they were collected If the
immediately after the disturbance may be signi- blocks have not yet been planted and no perma-
ficantly higher than before the disturbance. This nent reference points are available, the samples
nutrient flush is temporary, but still inflates the must be plotted as ciosely as possible on maps.
nutrient levels in the second samples.
6. Air-dry the samples and store them in a cool,
dry, environment until they are shipped to a
After the variation in soil types has been deter-
laboratory for analysis. DO NOT store soil
mined, soil fertility samples should be collected.
samples in plastic bags.
The variation in the soil types in each sampling area,
as determined by the soil pits, should be used to
Sampling for annual fertility assessment should
determine the minimum number of samples
be done in the fail e.g., September to November
required. NOTE: Chemical analyses of the upper
each year. Samples MUST be collected the same
horizon ( s) , the top 15 to 20 cm, from the soil pit
way each year, e.g., the depth to which the sample is
samples can also be used to supplement the cores.
collected will influence the nutrient levels .
It is important to collect samples distributed
Assessing Soil Fertility
throughout the sampling area. The following pro-
cedure is designed to ensure that the entire area is
Armson (1973) lists three objectives of soil and
sampled and that sufficient sample is collected. The
plant tissue analysis which apply to monitoring seed
major costs arise from processing and analyzing the
orchard fertility.
samples, NOT from collecting them. DO NOT CUT
CORNERS WHEN COLLECTING SOIL SAMPLES. 1. To determine why a tree exhibits poor growth
and/ororgan abnormalities such as discoloration.
1. Previously unsampled sites, sites with known 2. To detect nutrient deficiencies that inhibit
high variability e.g., seedling seed orchards on growth in a tree or stand.
cutovers: A minimum of 10 to 15 cores per 3. To control and regulate the nutrient supply to a
hectare should be collected. Sampling in a grid tree necessary to produce a crop to meet spe-
pattern at a 25 X 25 m spacing equals 16 samples cific objectives of management.
per hectare.
Previously sampled sites, and existing orchards In seed orchards, it is unacceptable to wait until
with the top 15 to 20 cm plowed and mixed to .
deficiency symptoms appear Nutrient deficiencies
create a homogeneous plow layer e.g., clonal that might inhibit tree growth and development
orchards: A minimum of 5 to 10 cores per must be detected before growth is seriously affected.
hectare should be collected. Sampling at 40 m X Therefore, the third objective given by Armson
40 m equals 6 samples per hectare. (1973) is the most important.
5-3
Soil acidity ( pH), should range between 5.0 and The organic matter content influences the
6.0 (Tables 5-1 and 5-2). At soil pH levels below 5.0, chemical characteristics and moisture holding
phosphorus availability decreases because of the capacity of the soil . Soils low in organic matter are
formation of insoluble complexes with iron and more susceptible to compaction by machinery than
aluminum ( Armson 1979) . The more soluble or soils with high organic matter content. The mini-
available forms of P occur in soils of pH from 5.5 to mum level of soil organic matter in the orchard
7.0. NOTE: Soil pH alone does not determine the should be between 4 and 5% (Tables 5-1 and 5-2) .
lime requirements. The clay and organic matter
content of the soil determines its buffering capacity
and hence the amount of lime required to change
the pH ( see APPENDIX VI).
Table 5-1. Suggested soil fertility levels for seed orchards, Canadian Forestry Service laboratory
Equivalent
extractable
Recommended nutrient
Units level1 (kg/ha)
pH 5.0 - 6.0
Cation
exchange meq /100g 10 - 15
capacity
P Available P -
75 100 450 - 600
( ppm) ( P205)
Table 5 -2. Suggested soil fertility levels for seed Cation exchange capacity
orchards. Nova Scotia Agricultural Col-
lege laboratory The cation exchange capacity ( CEC) of a soil is
its capacity to hold and exchange positively
Recommended charged particles ( cations) which include the nutri-
ents potassium ( K +) , calcium ( Ca 2 ) , magnesium
t
Units level 1
+
(Mg2+) , ammonium (NH/) , hydrogen (H ) and sodium
pH 5.0 - 6.0 ( Na+). The CEC of a soil is determined primarily
from its clay and organic matter contents and
organic (%) minimum 5.0 should be between 10 and 15 meq / 100 g.
matter
Soils with a low CEC have a low water holding
cation capacity, and lose N and K to leaching more readily
exchange than soils with a high CEC.
capacity meq/100 g
Soil macronutrients
N Total N
(%) The soil macronutrients include N , P, K , Ca and
Mg. They are required in relatively large quantities
P Extractable P 340 - 460 by trees. Fertility recommendations for seed
(P205) ( kg/ha) orchards in the Maritimes are based on tree growth
and fertilization studies. The optimum soil and
K Exchangeable K 280 - 380 foliage nutrient levels for cone and seed produc -
( KfeO) ( kg / ha ) tion are not known but can be estimated.
Table 5-3. Foliage sampling for nutrient analysis. The suggested numbers of shoots to be collected is the
minimum for the Canadian Forestry Service laboratory, and should provide ample material for
other laboratories
Table 5-4. Suggested foliar nutrient levels by species for seed orchards ( from Mahendrappa unpubl. data).
Orchard trees can be considered as being adequately supplied with the major elements if foliar
levels are maintained within these ranges
Species N P K Ca Mg
Black spruce 1.2 - 1.6 0.15 - 0.20 0.70 - 0.80 0.20 - 0.30 0.10 - 0.15
Red spruce 1.3 - 1.7 0.15 - 0.20 0, 60 - 0.80 0.20 - 0.30 0.06 - 0.10
White spruce 1.3 - 1.7 0.15 - 0.25 0.70 - 0.80 0.20 - 0.30 0.06 - 0.10
Jack pine 1.2 - 1.6 0.15 - 0.20 0.60 - 0.80 0.20 - 0.30 -
0.06 0.10
Red pine 1.2 - 1.6 0.20 - 0.25 0.60 - 0.80 0.20 - 0.30 -
0.10 0.20
White pine 1.7 - 2.0 0.20 - 0.25 0.70 - 0.80 0.20 - 0.30 0.10 - 0.20
Table 5-5. Suggested schedule for fertilizing individual grafts in seed orchards
NOTES: Fertilizer should be applied in a band (evenly distributed) around each graft. The band should be
located approximately half-way between the crown dripiine and the stem or, when applicable, at the outside
edge of the mulch. Do not DUMP the fertilizer at the base of the grafts. This can result in root damage and
possibly kill small grafts!
If heavy spring rains follow the early May application, it may be desirable to refertilize the grafts at the same
rates ( heavy leaching losses) . If this is done, the second scheduled application (mid-June) should be made
one month following the refertilization.
Literature Cited
CHAPTER 6
PEST MANAGEMENT
Seed orchards must be protected from damag- attributable to insects and diseases. Computer cod-
ing agents. Potential seed losses can be minimized ing forms for this system are found in APPENDIX
by isolating orchards from forests that provide VIII.
natural habitats for harmful insects, birds, mam- 1. Randomly select 10 ramets from each of five
mals, and alternate hosts of fungi. Adopting routine clones (10 seedlings from five families) foreach
cultural practices such as close mowing of the cover species and / or suborchard. These 50 trees
crop ( reduced small rodent habitat ) , and removing should be permanently tagged as they will be
old cones (several cone and seed insects overwinter used each year whenever possible. However ,
in old cones) will also afford some protection. All some replacement trees ( of the same
pest management operations must be recorded family/clone) may be necessary as not all trees
( APPENDIX IV ) . However, regardless of cultural will bear strobili every year.
practices, pests will invade the orchard.
2 . From early May to early June, preferably
BEFORE pollination, depending on species,
Assessing Cone and Seed Losses
count the total number of female strobili on the
sample trees . For large trees, tag two branches
An effective means of quantifying the impact or
and count the strobili on them. The tagged
potential impact of pests on a cone crop and
branches should be in the middle portion of the
concomitantly determining if a control program is
female cone -bearing region of the crown, and
warranted can be done through developing cone life
should have AT LEAST 20 strobili each. Tag as
tables. After the causes of losses have been IDEN-
many branches as necessary to obtain a total of
TIFIED and QUANTIFIED control efforts can be 40 strobili.
efficiently and cost-effectively undertaken. Gener-
ally , five steps are involved for each species in an 3. About 1 to 2 weeks following pollination, use a
orchard . hand lens to examine 10 strobili collected from
each tree for the presence of eggs or larvae.
1. Select sample trees . Many insects that feed directly on cones and
2. Obtain strobilus counts. seeds lay eggs during the pollination
3. Periodically monitor strobilus/ cone devel- period. If larvae or eggs are found, immediately
opment and quantify the losses over the grow- send a sample to the Canadian Forestry Service
ing season . - Maritimes, Forest Insect and Disease Survey
4. Determine actual seed yields from those cones ( F.i. D.S) for identification ( see APPENDIX VII) .
at the end of the growing season (at the end of 4. In August /September count the mature cones
the second year for pines). on the sample trees/branches. For pines, cone-
5. Compare actual yields with expected yields to lets are counted in the fall of year one and
determine the percentage of the potential mature cones the fall of the second year.
obtained. 5. Collect 10 mature cones from each sample tree.
Do a cut - test or extract the seed from these
APPENDIX VIII has detailed instructions on cones (see CHAPTER 7 for information on cone
developing cone life tables. maturation and assessing seed quality).
6. Calculate the seed yield and compare these
The following system for monitoring cone and values with the expected seed yield ( see
seed production is used to assess all aspects of seed APPENDIX VIII).
production efficiency, not just the losses directly
6 -2
There are currently NO insecticides registered Spruce cone rust ( Chrysomyxa pyrolae D.C.),
for the control of cone and seed insects in forest tree which can pose a problem for all spruce species, is
seed orchards in the Maritimes. Dimethoate, for use best controlled by eradicating the alternate hosts,
on Douglas-fir, is presently the only chemical regis- plants in the wintergreen family [ Pyrola spp. and
tered in Canada, specifically for the control of cone Moneses spp.) , within the vicinity of the orchard.
and seed insects in seed orchards. However , several Summers etal. (1986) obtained adequate control of
experiments testing the efficacy of different chemi- a western cone rust ( C. pirolata ) in a white spruce
cals in controlling cone and seed insects of Maritime seed orchard using the fungicide Ferbam. Ferbam
tree species have been conducted. Results from 76 WDG was recently registered for use against
these and other trials indicate that most of the pests spruce cone rust. One or two treatments applied
thus far encountered in Maritime orchards can be between one week before pollination and the end of
effectively controlled by one or more insecticides. pollination, provided effective control and increased
The major insect pests encountered to date in seed production in years of severe disease. How-
Maritime seed orchards and, where possible, ever, there were some indications that the fungicide
‘ promising ’ control measures are listed in affected seed quality. Further studies on the effects
APPENDIX IX. of Ferbam on seed viability are needed.
These trials also indicate that if cone and seed Controlling Other Pests
insects in Maritime seed orchards are to be control-
led effectively , orchard managers will require better Mammals and birds
tools than are currently available and better know-
ledge of the use of these tools. If these tools are to Mammals and birds can also cause consider-
become available, registration must be obtained for able damage to orchard trees. Voles and field mice,
the desired chemical (s). The demand for chemicals which girdle young trees , may be effectively
for orchard use is not likely to be sufficient to controlled by close mowing of the cover crop and
persuade most chemical manufacturers to conduct other vegetation (especially in the fall) , but toxic
the research required for full registration. The most baits are still the most effective method of control-
viable alternative is for Minor Use Registration. Taky ling field mice ( Peterson 1982). Examine the orchard
(1986) provides details on Minor Use of Pesticides in September or October for signs of mouse activity.
Program in Canada ( copies of this publication are Ontario Ministry of Agriculture and Food (1986) lists
available from Agriculture Canada ). Orchard some of the mouse baits that are available as well as
managers in the Maritimes must make a coordina- their application rates:
ted effort to ensure that some of the chemicals with 1. Waxed zinc phosphide bait- 5.5 to 11.0 kg /ha. If
potential use in seed orchards are registered under further mouse activity detected, repeat when
this minor use program. rain is not expected.
2. Ramik Brown- 22 kg / ha in two applications of 11
Control of diseases kg with 20 to 40 days separating treatments.
3. Rozol paraffinized pellets ( 0.005 % chloro-
Diseases may also affect orchards. Disease phacinose) - 11 kg/ ha .
problems will be reduced by carefully selecting a
proper site and practicing good sanitation To minimize the danger of poisoning nontarget
techniques . animals such as birds and dogs, establish a series of
feeding stations around the orchard. The bait can be
Needle rusts ( Pucciniastrum spp.) may also placed in several types of containers including T-
pose a problem in orchards. They can cause defolia- shaped pieces of PVC pipe, ice cream containers
tion and subsequent growth loss, however, they do with holes cut in them, or styrofoam cups with holes
not usually kill large trees. Needle rusts can cause ( two cups with their tops taped together ). Stations
considerable damage to cones and seeds ( see should be concentrated around the perimeter of the
Smith et al. 1986) . To date, the use of fungicides in orchard ( reducing influx of mice) and several
seed orchards to control needle rusts has not been stations should be located within the orchard to
required. control existing populations:
6- 4
AcJeris valiana
Adelges abietis
Adelges lanciatus
82
Adelges piceae
Adelges strobilobkis
Aphrophora spp.
Asynapta hopkinsi
Barbara mappana
Choristoneura fumiferana
Choristoneura pinus
Cmara spp
Goleophora laricelia
Cofeotechnites I arid s
Corjopftffton/s banksianae
-
4
3BB8W
Conophthorus coniperda
Conophthorus resinosae
Cydia si rob HeIla
Cydia toreuta
Dasineura canadensis
Dasineura rachiphaga
Dendroctonus rufipennis
Dendroctonus simplex m
Dtoryctria abietivorella
Dioiyctria disclusa
Dioryctria renlculelloldes
Ectropis crepuscularia
Endoptza piceana
Eucosma monitorana
Eucosma tocullionana
Eupithecia albicapitata
Eupithecia. mutata
Exoteleia nepheos
Formica spp.
Gif pin !a hercyniae
Henricus fuscodorsanus
Holcocerina immaculella
Figure 6-1. Major insect pests and their feeding times in Maritimes tree seed orchards (from Forest Insect and
Disease Survey , 1986., unpublished data)
6- 5
Hylemya anthracina
Hylemya viarium
Hylobius spp ,
Lambdina fi seel lari a fi seel laria
Mayetiofa carpophaga
Mayetiola piceae
Neodipnon swainei
Neodi prion Virginian a
Ofigonychus milled
Ofigonychus ununguis BOB
Orgyia leucostigma
Petrova albicapitana
Physokermes piceae
Pikonema alaskensis
•''• ••*'**'*'
W i 1 I
Pissodes strobi
Pissodes spp.
Pleroneura brunneicornis
Pristiphora erichsonii
Resseliella spp,
Rhabdophaga swainei
Rhyacionia buoliana
Spilonota lariciana
Tetyra bipunctata
Toumeyella parvicornis
Xyela spp.
Zeiraphera canadensis
Zeiraphera improbana na
6 -6
Snowshoe hare and porcupines, which chew when there is a person residing at the orchard year -
bark off trees and squirrels which cut cones, can be round. However, for most orchards, this is not
controlled by placing metal bands around the stem .
practical Placing signs around the perimeter of the
of each tree. To be effective, bands must extend up orchard indicating that the area is a seed orchard
the tree higher than the hares or porcupines can may act as a deterrent. Public education, including
reach when standing on deep snow. Banding is tours of the facility for local residents may be most
expensive and is NOT practical in young seedling beneficial. Employing a trusted and respected local
seed orchards ( 2500 stems / ha) before final rogue- resident to conduct periodic checks may also be
ing or for small grafts. Small trees can be protected worthwhile.
by surrounding them with chicken wire, hardware
.
cloth, or plastic ‘sleeves' Fire
Access to seed orchards by tree poachers, Above-tree mist -irrigation systems are effective
vandals, all-terrain- vehicles and snowmobiles is in reducing frost damage. Other means of reducing
difficult to control. The best control can be obtained frost damage include distributing heaters through -
in those orchards which are completely fenced and out the orchard and fanning the morning frosts with
6- 7
helicopters. All frost protection methods are expen - Miller, G.E. 1983. When is controlling cone and seed
sive, but the potential ramification of doing nothing insects in Douglas- fir seed orchards justified?
is to lose entire cone crops. For. Chron. 59:304-307.
CHAPTER 7
SUMMER i SUMMER
r
V? (S'feC
Uyn
.
;••>•/ » Tc'Wlrr
(
/ ytr; .!
* ''
/ Cwn /
/
Co -
'- i’S I
r;
: . * .>#< 0>
Ca< cS \ -
*>V F
SPRING C&nn
- ,
FtrtBe ta
X FAU •
SPRING
FWhj S
iXCilfV
\
X PALL
* / \
/ \
/ \ \
/ \ \
/
/ \ s \
/ \ / \
/ \ / \
INTER i ; WINTER;
•* *
Figure 7-1. The reproductive cycle of white spruce Figure 7-2. The reproductive cycle of jack pine
(adapted from Table 7-1). (adapted from Table 7-1).
7-2
Estimates of times or periods of initiation and Limited research has been conducted into cone
differentiation are given in Table 7-1 for average and seed enhancement forspecies in the Maritimes.
years in the Maritimes . A late or early spring will The recommendations given are based largely on
alter these times by as much as two weeks. Warm, trials with black spruce in New Brunswick , white
southern locations tend to have earlier dates for spruce in Nova Scotia, and the remainder are from
initiation than cool, northern locations. Similarly , reports in the literature. A detailed review of the
favorable aspects ( southerly) elicit earlier develop- literature on many aspects of cone production in
ment than less favorable aspects ( northerly). The forest trees is available in Owens and Blake (1985) .
times of differentiation are less diverse than those of
initiation. For the spruces, initiation occurs during
the 10 days before vegetative buds burst, and
differentiation occurs at or just after shoot elonga-
tion ceases.
Table 7-1. Estimates of times of periods of initiation of potentially reproductive buds, and of differentiation of
reproductive structures within buds, forspecies growing in average conditions in the Maritime
Provinces ( Powell 1983 with modifications by personnal communications 1986)
Species Initiation Differentiation
’Provisional dates. Investigations are continuing (Powell personal communication Dec . 1987) .
7-3
Fertilizers Weather
6" Stake r \y
/
Wooden bearing
on t o p )
^-
( I / 2" dowel tapered
The numbers of pollen and seed cones must be
evaluated in the spring ( strobilus counts) see
APPENDIX X and again in late summer/fall ( seed
quality assessment). The same trees used for monitor-
ing cone development should be used e.g. , 10
Figure 7-3. Diagram of a pollen trap (from ramets from each of five clones or 10 seedlings from
Greenwood and Rucker 1985 ). five families (see CHAPTERS 5 and 6) .
Table 7-2. Evaluating cone crop seed yields from a The number of cones per tree and number of
cut -test seeds per cone are used to estimate the total
expected seed yield from the orchard, and to com-
Species Number of full seed per half cone pare estimated seed yields with those actually
obtained (see next section) .
Low Medium High
When orchards are young, seed production
Black spruce <6 6 -9 > 10 varies considerably between trees, often with most
Red spruce <6 -
6 9 > 10 of the seed being produced by a small number of
White spruce <6 6-9 > 10 clones or families. Also, since little within- orchard
pollen may be produced , the genetic quality of the
Jack pine1 <10 10-15 > 16 seed may be low because of pollen contamination
Red pine < 6 6-9 > 10 and/ or self -pollination, or seed-set may be low
White pine <10 10-15 > 16 because the orchard is well isolated from contami-
nating pollen. Although the quality of seed from
Tamarack < 2 -
24 > 5 such crops may be low, they must be collected to
reduce habitat for cone and seed insects.
jack pine cones is difficult so it might be
’Slicing
preferable to extract the seed. Time of collection
White pine
Tamarack IIIIIIIIIIHIliillBIBBBB
tllllllllllllllll Embryo maturation : too early to collect cones , ie. , seed will probably not
complete development regardless of subsequent cone handling .
|& HS 0 ES9 63 J23 Cone ripening : cones can be collected but will require after - ripening
before seed is extracted.
Cone collecting : cones are mature and can be collected .
Figure 7- 4. Cone and seed maturation stages for eight Maritime tree species. Development stages can be
advanced 1 to 3 weeks with warm and dry weather and sites, and similarly retarded on cold and
wet sites or where growing seasons are naturally later e.g., the Fundy shore (from Smith 1985 ).
1981;1983). Correlations of seed maturity with ficult to remove by twisting and pulling, while white
degree day accumulations should be determined and red spruce cones are easily removed by the
for each orchard and each orchard species since latter method.
seed will probably be shed earlier from orchard
trees than from trees in mature stands. When trees are short , ladders, particularly
tripods can be efficiently used to collect cones
How to collect ( Yeatman and Nieman 1978). As the trees grow, and
access to cone producing parts of the crown is more
Cone collection crews must receive proper difficult, hydraulic platforms and moveable
training in collection techniques ( see Dobbs et al. scaffolds ( scaffolds mounted on trucks ) are
1976). Seed loss cannot be tolerated. Only cones necessary. 'Cherry-pickers’ or bucket booms offer
and not shoots should be removed, because a shoot good maneuverability around tree crowns but are
that bears seed cones one year, will eventually expensive to purchase and can compact the soil.
produce other shoots which will bear seed and Hallman and Casavan (1979) evaluate much of the
pollen cones. Ease of removing cones from the equipment currently available for collecting cones.
shoots differs between species. Jack pine cones Most of the equipment reviewed has not been tested
should be cut off with clippers because they are in the Maritimes.
firmly attached by a stout stem making them dif-
7- 8
Cone handling is an extremely important step in Dobbs, R .C.; Edwards, D.E.W.; Konishi, J.; Wallinger,
the cone collecting process. Cones must not be D. 1976. Guidelines to collecting cones of B.C.
collected and stored ‘ wet ’. They should be packed .
conifers British Columbia For. Serv./ Can. For .
loosely in half filled burlap bags, placed in a well Serv., Joint Rep. No. 3.
ventilated garage or cone- shed, and shipped as
soon as possible to the extractory . Label each bag Ebeli, L.F. 1972. Cone induction response of
inside and outside according to species, orchard, Douglas-fir to form of nitrogen fertilizer and time
sub -orchard or orchard block , date collected, and of treatment. Can. J . For. Res. 2:317-326.
any other necessary information ( see Smith 1985).
The orchard managers’ responsibility and interest Fashler, A.; Devitt , W.J.B. 1980. A practical solution
in the seed should not stop when the cones are to Douglas-firseed orchard pollen contamination.
shipped to the extractory. The manager should For. Chron. 56: 237-240.
ensure that:
1. The cones are stored properly at the extractory. Fraser , D. A . 1975. Management of tree growth and
research plantations. Pages 192-201 In Proc. 12th
2. The extraction of the seed is both safe and
Lake States Tree Improv. Conf. USDA For.
efficient.
Serv. , Gen, Tech. Rep. NC-26.
3. The seed yield information and results from any
seed tests that have ( or should have) been Friedman, S.T.; Adams, W.T. 1981 . Genetic effic-
conducted are received. iency in loblolly pine seed orchards. Pages 213-
224 In Proc. 16 th South . Forest Tree Improv.
This information combined with the other Conf., Virginia Polytech. Instit. State Univ., May
orchard records is necessary to assess orchard
productivity accurately and to compare on -site
27-28, 1981 .
estimates of seed yields and quality with those Greenwood, M.S.; Rucker, T. 1985. Estimating pollen
reported from the extractory . Over the long term, if contamination in loblolly pine seed orchards by
these records are accurately maintained, the pollen trapping. Pages 179-186 In Proc. 18 th
experienced orchard manager will know if the cones South. For. Tree Improv. Conf ., Long Beach,
were handled properly after they left the orchard. Miss., May 21-23, 1985 ,
APPENDIX I
-
Figure I 2. Drainage
Sandy loam
^Shaltow
soil overlying
sandy > loamy sand
loa
White White
Spruce Pine
verify *
an map °
'
ral site P
gene
r,gure V 4 Orchard
'
11-1
APPENDIX II
ASSUMPTIONS
No. cones /graft: 200
Sound seed/ cone: 30
Seedling production method: container; 1.5 sound seed/seedling
Interval between cone crops: 2 years
Area required to
produce seed for
1 million seedlings = (1,000,000 seedlings) / (1,100,000 seedlings/ ha)
= 0.91 ha
Because of 2 year periodicity between cone crops, twice this area is required to produce enough seed for
1 million seedlings annually.
Total area required to produce
5 million seedlings/ yr = 1.82 ha x 5
= 9.1 ha
This area requirement is dependent on the assumptions. For example, if the average number of cones per
graft increases to 300 because of good management, then the orchard area requirement is reduced by
one-third or if 2 full seed are required to produce a seedling, the orchard area is increased by one-third.
IIM
APPENDIX III
Seed Orchard:
Site History
Site Preparation
2. method: Date(s):
Pre-sowing amendments:
Root -pruned trees are tempora'rily weakened seed orchards Fast growing species such as
and may be more susceptible to infection from root tamarack can grow a metre or more per year when
rots . When pruning, whether by discs or spades, be maintained under the high fertility regimes in seed
sure to fill in the line of disturbance. Exposed roots orchards. Such trees quickly become too tail for
increase the chance of attack by insects and disease. easy cone harvesting. Top-pruning should begin at
Trees that are repeatedly root -pruned may not be as an early age. For mostspecies in the Maritimes, this
wind - firm as unpruned trees and may be more will probably be between age 7 and 10 years,
susceptible to drought. whereas vigorous grafts of larch will require pruning
before age 7 . During the early years of orchard tree
Girdling and strangulation development trees can be topped regularly to pro-
mote large bushy crowns capable of carrying a large
Girdling by making cuts in the tree or branch number of cones. However, topping may not be
and strangulation by placing bands, usually metal, used in seedling seed orchards until they have been
on the tree to constrict growth can induce water .
rogued at least once The degree to which top-
stress and affect the movement of carbohydrates pruning can and should be used for Maritime tree
and nutrients in the tree. Results from girdling and species, needs to be further investigated.
strangulation experiments have been variable, but it
appears that, when timed properly, both can stimu - Pollen Contamination
.
late a ‘one-time’ heavy cone crop However, these
methods are not recommended in seed orchards. To maximize the genetic quality of the seed
Tree mortality can result if damage is too severe and produced in an orchard, pollen contamination must
long - term cone production may be less on treated be minimized. Studies in the southeastern United
trees than on control trees because of the overall States indicate pollen contamination can be as high
reduced vigor. as 30 -80%, even in mature orchards in which the
trees are producing large quantities of pollen. This
Growth hormones can reduce genetic gain by at least 2-4% ( Friedman
and Adams 1981 ) . Preliminary studies in several
In recentyears, trials with growth hormones, parti- Maritime orchards indicate background pollen
cularly gibberellins, have been successful in enhanc- levels are high.
ing both pollen and seed cones in many species.
This is particularly useful for stimulating individual Foreach orchard, establish a pollen monitoring
trees orclones to produce cones for use in breeding system to determine the amount of contaminating
programs. Three major drawbacks to operational pollen and to assess its potential impact on the
use of growth hormones in seed orchards are quality of seed. Pollen arriving either before or after
evident; the period of strobilus receptivity is of little
consequence.
1. The concentrations most effective in field trials
have sometimes resulted in problems with phyto- 1. Estimate background pollen levels by measur -
toxicity associated with the surfactant. ing pollen levels while the trees are young ,
2. Cost of the hormone is prohibitive. BEFORE they produce significant quantities of
pollen.
3. The techniques for large-scale applications are
not yet sufficiently refined. 2. Continue monitoring pollen as the orchard trees
mature. The difference between the two levels
However, hormones may be an important tool provides an estimate of within -orchard pollen
for seed orchard managers in the near future. production.
APPENDIX IV
Table IV -1 provides a form to record an annual summary of all operations conducted to aid the orchard
manager in planning year to year operations and time and manpower requirements. The remarks section
should contain general comments such as manpower used, time to complete work , area, block number,
number of trees treated.
Operations to Include
Establishment
planting
herbicide application
mulch application
graft pruning
Vegetation management
cleaning
herbicide application
Pest management
insect /disease monitoring
pesticide application
Fertility management
broadcast applications
individual tree applications
soil/ foiiage sampling
Cone crop management
monitoring/ forecasting
pollen monitoring
fertilizer application
root pruning
hormone application
top pruning
harvesting
Rogueing
Grass mowing
Year:
APPENDIX V
FERTILIZATION RECORDS
The Fertilization Record field sheet (Table V-1) can be used to record the day- to- day operations. This
information is then copied onto the Fertilizer Application Record form ( Table V-2) for computer entry. The
Orchard Fertility Record form ( Table V -3) , is used to record results from soil and foliage analyses. After the
data from these forms are entered in the computer, programs can be used to produce detailed fertility
assessments/reports. These records can be updated as required.
NOTE: Although there are two separate forms, (convenience of data input ), the data from both can be merged
and a comprehensive summary produced, e.g., monitor changes in soil fertility levels with fertilizer
applications.
V-2
Instructions for completing the Orchard Fertilizer Application Record - field sheet . All details regarding
fertilizer applications should be recorded. Each form contains the data for one block and may be used for one
or several years, as deemed necessary.
1. Broadcast applications
Application method: Type of equipment used e,g„ drop-type ( GANDY ) spreader vs. cyclone spreader.
1. Any 'problems’ with applying the fertilizer e.g., equipment calibration . Can also note any equipment
settings for future reference.
2. Weather e.g ., very heavy rains shortly after applying fertilizer (especially important for fertilizers, such
as ammonium nitrate, that are prone to heavy leaching losses) .
Fertilizer placement: e.g., band around the crown dripline, bands on two sides of the trees, etc.
Column
No.
others as required
BROADCAST APPLICATIONS
6-7 % N
8- 9 % P2O5
10-11 % K 2O
12-13 % Ca
14-15 % Mg
0- 46-0 triple-super-phosphate
etc.
16-21 Date:
16-17 Day
18-19 Month
20- 21 Year
38 - 43 Date: As above
44- 48 Rate: grams fertilizer per tree or if fertilizer was applied in a band, specify the
rate expressed as grams fertilizer per square metre, etc .
REMARKS
-
49 80 Note any factor that could affect the effectiveness of the fertilization operation
etc.
V- 8
Column
No.
N
. °
P P2 5
K, K 20
Ca, CaO
Mg, MgO
pH
Cation exchange capacity ( C.E.C.)
. .
Organic matter ( O M ) .
11-18 Units
21- 26 Date:
Date that the samples were collected.
21- 22 Day
23-24 Month
25- 26 Year
27- 32 Level: the nutrient level expressed in the units defined in cols. 11-18.
APPENDIX VI
Table VI-1. PH
Table VI-2. Phosphorus
Table VI -5 . Magnesium
No attempt was made to assemble a nitrogen table because soil nitrogen varies with time of collection,
organic matter content , etc.
VI- 2
Soil pH
texture
3.5-4.5 4.5-5.5 55-6.5
Table VI-2. Soil phosphorus. Available P 2O5 calculated for a kg -ha-15 cm.
Amount of
Available p o / triple super Amount of
2 5
P ha-15cm phospate re- P2O5 added
(ppm ) ( kg ) quired (kg/ha) (kg/ha) Comments
Amount of
Exchangeable K 2SO4 Amount of
K K 20/ha-15 cm required «20 added
meq/100 g ( kg) (kg / ha) (kg ha )
/ Comments
0.05 61.3 300 147 Very low. May require a 2nd appli-
0.10 122.6 300 147 cation. Do not apply more than 300
0.15 183.9 300 147 kg/ ha K 2 S04 in one year as K
0.20 245.2 250 123 leaches readily.
0.25 306.5 150 74
0.30 367.8 100 49 Low K .
0.35 429.2 50 25
0.40 490.5
0.45 551.8
0.50 613.0
-
Table VI-4. Soil calcium. Available CaO calculated for a kg ha-15cm.
Amount of Amount of
Exchangeable CaO dolomitic lime CaO
Ca ha-15cm required added
meq/100 g (kg) (kg/ha) (kg/ha) Comments
Note: TheCa:Mg ratio should not exceed 10: 1. Therefore the Caand Mg levels should be evaluated together.
The pH of the soil must also be considered before applying dolomitic lime.
VI - 4
Amount of Amount of
Exchangeable MgO dolomitic lime MgO
Mg ha-15cm required added
meq/100 g (kg) (kg/ha) ( kg/ha) Comments
Note: The Ca:Mg ratio should not exceed 10:1. Therefore the Ca and Mg levels should be evaluated together .
The pH of the soil must also be considered before applying dolomitic lime.
VII-1
APPENDIX VII
Fertilizers can be mixed in different ratios depending on how much nutrient you want to apply per given area.
These mixes will have the nutrients in the same proportions as commercial mixes. However, LESS total
material will have to be applied to obtain the same nutrient additions than were a commercial mix used.
Drawbacks arise because you need to purchase several different fertilizer types ( need to buy in bulk ) and you
.
require the equipment to mix the materials. THEY MUST BE MIXED THOROUGHLY!!!! e. g , in a large cement
mixer.
Examples of how several single-element fertilizers can be combined to produce a desired mix are given below.
Calculations
1. Start with the fertilizer with the nutrient which has the lowest percentage e.g., ammonium nitrate 34%
( versus 48% K 20 and 46% P2Og).
2. Calculate ratios of the nutrient percentages, i.e., smallest over each of the other two:
34/48 = 0.71
34/46 = 0.74
3. The proportions ( by weight ) of the three fertilizers which must be added to obtain a balanced mix, e.g.
10-10-10, would then be
Example I
A 10-10-10 fertilizer contai ns 10% N, 10% K 20, and 10% P 205 and if you wanted to apply the equ ivalent of 500
kg/ha of 10-10-10
500 kg X 0.10 = 50 kg N
50/0.34 = 147 kg ammonium nitrate required to add the same total amount of elemental nitrogen ( N) .
From the practical standpoint the quantities would be rounded. You would only be applying 360 kg fertilizer
mix whereas with the premixed 10-10-10 you would have to apply 500 kg.
Example II
46/ 46 = 1.00
46/ 48 = 0.96
46/ 46 = 1.00
Ratios
urea 1.00
potasium sulphate 0.96
triplesuperphosphate 1,00
500 X 0.10 - 50 kg N
50/.46 - 109 kg urea required to add the same total amount of elemental nitrogen (N).
-
1,00 X 109 109 kg urea
0.96 X 109 - 105 kg potassium sulphate
-
1.00 X 109 109 kg triplesuperphosphate
You would apply only 323 kg fertilizer mix whereas with the 10-10-10 premix you would apply 500 kg.
Note: For all practical purposes, the percent nutrient content for these three are close enough that they can
be mixed equally and produce a balanced mix .
VIII-1
APPENDIX VIII
Strobilus and cone pest damage assessments, and instructions for constructing cone life tables. The
assessment codes for insects and diseases include most pest species directly attacking cones and seed of
Maritime conifer species.
Column
No,
1- 21 TREE IDENTIFICATION
1- 2 Orchard Number: Agencies with several different orchard sites or suborchards within a single
complex may wish to assign a 2-digit code to distinguish them.
Lt Larch, tamarack
Pj Pine, jack
Pr Pine red
Pw Pine, white
Sb Spruce, black
Sn Spruce, Norway
Sr Spruce, red
Sw Spruce, white
5-7 Block
8- 10 Row These entries are used to locate EXACTLY which tree was sampled. This
makes identifying the same tree easier .
11-13 Column
If desired, more intensive sampling may be done, e.g., conelet tagged individually and its position
noted.
0101 whorl 1 conelet #1
0102 whorl 1 conelet #2
0201 whorl 2 conelet #1
0202 whorl 2 conelet #2
This system applies more to a research study than to operational monitoring. However, the results;
can be used for operations as well.
26 Branch number: If individual branches are tagged, a separate number is used for each.
27- 66 DAMAGE CODE
27- 28 Pest: Number code to distinguish the type of damage ( if any) observed. See Table Vlli-1 for a
suggested coding system.
29-30 Species: A combined letter-number code to identify the pest species ( Table VIII-2) .
.
e.g , A1 Adelges abietis
If more than one pest is found in one cone, then each should (can ) be listed separately and noted in
the comments section. The number of healthy cones is the check to ensure the total is not changed.
31 ID: In many instances, the orchard manager will not be able to identify the insect species on the
.
samples When this occurs the sample should be shipped to the local FIDS office for identification.
The ID code can be used to indicate whether or not a sample was sent to FIDS.
Y - Yes
N - No
A copy of the form for submitting samples to FIDS is given in Fig, VIII-2.
32- 36 Date sampled
37- 51 as per 22-36 above,
52- 66 as per 22-36 above.
67-80 COMMENTS
General notes on developmental abnormalities, etc. Other cross-reference notes should be made
here (e.g., branch damaged after sampling). Agencies can develop their own coding system for
comments, e.g.,
brdam branch damaged after sampling
VIII-3
02 Diseases
06 Unknown
INSECTS
Species
code Scientific name Common name
Species
code Scientific name Common name
DISEASES ( FUNGI)
!
VIII- 9
Figure VIII- 2. Seed orchard Forest Insect and Disease Survey (F.I.D.S.) sample submittal form.
VIII- 10
Telephone: Collector:
Seed Orchard
Host (species)
Type of sample (no.) Soil
Date Collected
Foliage _ Cone
Stock Type: graft rooted cutting _ seedling
other (describe)
Number of ramets /clone or seedlings/ family sampled
Describe how the samples were collected: e.g. for soils-state whether composities were made or just
single cores, depth of soil sample; for foliage-where in the crown foliage was collected, etc...
NOTE: It is recommended that both soil and foliage samples be collected when nutrition, or other abiotic
problems are suspected.
C. PROBLEM:
CONDITION: Describe problem: include information on first appearance, present symptoms, rate of
spread, mortality, unusual problems (such as soil nutrient, water or climate), etc. Be as complete as
possible, include helpful “hints”.
D. DELIVERY: Include submittal form, mark package PERISHABLE and send to: Forest Insect and
Disease Survey, Canadian Forestry Service - Maritimes, P.O. Box 4000, Fredericton, N. B. E3B 5 P7.
Instructions for completing Figure VIII -3 Seed Yield form, and examples of cone life tables based
on the
procedures given in Bramlett and God bee (1982) . The data from Figures VI 11 -1 and VIII-3 can be combined in
the computer and all necessary calculations performed.
Column
No.
1- 2 Orchard Number: Agencies with several different orchards sites or sub-orchards within a
single complex may wish to assign a 2-digit code to distinguish them.
Lt Larch, tamarack
Pj Pine, jack
Pr Pine red
Pw Pine, white
Sb Spruce, black
Sn Spruce, Norway
Sr Spruce , red
Sw Spruce, white
5-7 Block These entries are used to locate EXACTLY which tree was sampled. They
make returning to the same tree easier.
8-10 Row
11-13 Column
22- 24 Number of strobili (NS ): Number of strobili counted on the sample tree.
25- 27 Number of branches (NB ): Number of branches on the sample tree on which strobili
were counted.
28-30 Total number of branches (TNB): Predicted total number of cone-bearing branches on the
sample tree.
VIII- 12
31- 34 Predicted total number of strobili ( PTNS): If all the strobili on a given sample tree were
counted, then
PTNS = NS,
otherwise
PTNS = NS x TNB/NB.
35- 38 Sound seed ( SS ): Number of sound seed obtained from the cones. Either one line per single
cone or a total for the cones (see 39- 41 below) .
39- 41 Cone number ( CN): This is the number of cones used to obtain the sound seed yield data, e.g,
not all the cones on the sample trees will be sampled and have their seed extracted.
42- 45 Seed potential per cone ( SPPC): This value is the number of fertile cone scales times two, e.g.,
the biological potential per cone. Values for this should be determined for each species and
orchard by counting the number of cone scales in the middle two- thirds of the cone. Few good
seed is obtained from the scales at the top and the bottom of cones. These values should not
vary greatly between clones or families nor between years within a given orchard.
If this information is not obtained, or not known for a given orchard, the following approximate
values can be used.
46- 49 Predicted sound seed per cone (PSSC): The PSSC values are determined based on past seed
yield data . For the first sampling year,
PSSC = SS/CN
50- 53 Predicted total number of seeds per tree ( PSPT): Once the total number of strobili on the
sample trees has been determined, a PSPT value is calculated:
54-57 Number of trees ( NT ): This is the total number of ramets or seedlings in the orchard or
orchard block .
58- 61 Cone size ( CS): Number of cones per litre. This value should be determined for each orchard.
Cone size will vary with parent tree, species, and orchard site ( fertility ) .
62-65 Predicted orchard seed production ( POSP): The predicted orchard seed production ( POSP) ,
expressed in thousands of seeds is the cumulative total of all the sample trees. The total count
( PSPT ) is averaged for all the sample trees and then multiplied by the total number of
seedlings /ramets in the orchard.
e. g.,
where:
66- 69 Predicted cone efficiency ( PCE): The predicted cone efficiency is an expected value for a
given years ’s actual cone efficiency ( ACE). It is not measured but is the best estimate available
based on knowledge of past orchard performance.
We do not as yet have accurate ACE values for Maritime orchards . Cone efficiency values from
the southern U.S. ( Bramlett and Godbee 1982) are as follows:
Accurate estimates for ACE will be obtainable only after cone efficiency values have been
measured in an orchard forseveral years. Until thattime, the values from Bramlettand Godbee
(1982) can be used.
VJII- 14
70- 73 Predicted seed efficiency (PSE ): Predicted seed efficiency is the ratio of filled seeds (SS )
to the seed potential ( PSSC) . A similar procedure to calculating PCE is followed.
For the first year of inventory, a "best estimate" should also be used. Again, as there are
no values for Maritime seed orchards, suggested estimates are taken from Bramlett and
Godbee (1982 ).
For subsequent years, PSE values should be obtained from the average of previous years.
74 -76 Estimated extraction efficiency (EEE ): The estimated extraction efficiency measures the
percentage of the sound seed removed from the cones.
Orchard predicted extracted seed ( OPES ) ; The OPES is based on the results from previous
cone analyses and is calculated as follows:
77- 80 Seed use efficiency (ESU ); The ESU is a measure of reductions from the total seed potential
which occur at the nursery. The two main sources are;
77-78 1. Excess seed usage (ESU):i.e., two seed sown per cavity without transplanting the extra seedlings.
79- 80 2. Reduced germination ( RG ): It is not reasonable to expect 100% germination, hence this
figure will be less than 1.0.
ESU * 100/ESU X RG
e.g., two seeds per cavity; ESU = 2
90% germination; RGE = 0.90
The data from these procedures must be collected accurately and regularly if the productivity of an orchard
is to be correctly assessed. When cone life tables (Figure VIII-4 ) have been constructed , measures can be
taken to identify which sources of seed loss are most important, and consequently where to direct
preventative measures to best rectify the problem ( s ) .
VIII-16
TREE IDENTIFICATION
a: DATA SEED PRODUCTION ESTIMATING EFFICIENCY
oo <
CLONE / LJ ON TOTAL SEED
PILLI PER TREE
STROBILI
FAMILY >-
LU
< CDo O z> PRODUCTION P
X o $ LLI P E SUE
O 2 LLJ o J
C S
o NUMBER Z E
gl co CL
CD r
< o o NS
o
NB TNB PTNS SS CN SPPC PSSC PSPT NT CS POSP E E E ESU RG
2 3 4 5 6 7 8 9 I0 M I 2 I3 |4 15 16 17 18 19 202 ! 22 2324 25262 ? 2E 29 30 31 32 33 34 35 36 37 38 3 E 4 C 4 ! 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 5E 59 60 61 62 63 64 65 66 67 68 69 7 C 71 7273 74 75767776 7 80
‘
vm-17
100 5%
o,
V20%
Q.
2 80-
o
a;
c
5 60-
o
' o White spruce
A
§ 40-
<v
QL
20-
100
80-
C
a> 80
o
a)
CL
60-
“O
a>
40
a) 54
cn 5I
o
c
40- B
CD
o
CL
20-
80 % 75 % 90 % 95 %
Cone Seed Extraction Seed Usage
Efficiency Efficiency Efficiency Efficiency
( PCE ) ( PSE ) ( EEE ) ( SUE )
Figure VIII- 4. Cone life- tables A - cone efficiency ( survival) , and B - seed orchard to nursery efficiency
(adapted from Bramlett and Godbee 1982).
XI- 1
APPENDIX XI
REPRODUCTION OF CONIFERS
APPENDIX IX
Taky (1986 ) 1 lists some insecticides and their formulations which are registered in Canada which have
.
potential utility in seed orchards under the pesticide Minor Use Program As most are NOT registered
specifically for use in seed orchards, refer to the compendium of registered pesticides ( Agric . Can . 1984) 2 for
the exact restrictions placed on these chemicals.
The following criteria must be met before a proposal for minor use will be entertained:
1. The pesticide must have been evaluated and registered for other purposes under the Pest Control
Products Act.
2. The use must be for a crop or pest for which adequate pesticides are not already registered.
3. There must be adequate reasons, or experience, to believe that the pesticide will be effective for the
expressed, intended purpose.
All of the chemicals listed in this section meet these criteria for the target insects given. Recommendations are
not given for all insect species on the list ( see tables IX-1, IX-2) .
1Taky, J. 1986. Minor use of pesticides program handbook. Pesticide Information. Special Edition. Agric.
Can., Res. Branch. Ottawa.
Agriculture Canada. 1984. Compendium of pest control products registered in Canada: Control of
arthropods and molluscs. Agric. Can. Pesticides Division, Plant Health and Plant Products Directorate.
1984.
IX - 2
Table IX -1. Insect species listing cross-referenced with ‘potential’ control measures
Code Species Control Codes ( see table IX-2)
A1 Acleris variana
A2 Adelges abietis IV,XII, XV,XVI
A3 Adelges lariciatus
A4 Adelges piceae IV, XV
A5 Adelges strobilobius
A6 Aphrophora spp.
A7 Asynapta hopkinsi
B1 Barbara mappana
CO Choristoneura fumiferana . .
VI VII XII
C1 Choristoneura pinus
C2 Cinara spp. I,IV,V,XII
C3 Coleophora laricella IV, XII,XVI,XVII
C4 Coleotechnites laricis
C5 Conophthorus banksianae
C6 Conophthorus coniperda
C7 Conophthorus resinosae
C8 Cydia strobilella
C9 Cydia toreuta
D1 Dasineura canadensis
D2 Dasineura rachiphaga
D3 Dendroctonus rudpennis
D4 Dendroctonus simplex
D5 Dioryctria abietivorella
D6 Dioryctria disclusa
D7 Dioryctria reniculelloides
E1 Ectropis crepuscularia
E2 Endopzia piceana
E3 Eucosma monitorana
E4 Eucosma tocullionana
E5 Eupithecia albicapitata
E6 Eupithecia mutata
E7 Exoteleia nepheos
G1 Gilpinia hercyniae
HI Henricus fuscodorsanus
H2 Holcocerina immaculella
H3 Hylemya anthracina
H4 Hylemya viarium
H5 Hylobius spp.
M1 Mayetiola carpophaga
M2 Mayetiola piceae
M3 Megastigmus atedius atedius
M4 Megastigmus lands
M5 Megastigmus specularis
M6 Mindarus abietinus
N1 Neodiprion abietis
N2 Neodiprion nanulus nanulus .
IV, XII, XIII, XIV, XVI, XVII
N3 Neodiprion sertifer
N4 Neodiprion swainei IV,XII , XVI
N5 Neodiprion virginiana
01 Oligonychus milleri
02 Oligonychus ununguis I,IV,V,VIII,IX,X, XII
03 Orgyia leucostigma XII, VII
P1 Petrova albicapitana
P2 Physokermes piceae IV , VI, XII
P3 Pikonema alaskensis XII,XIII,XIV , XVI, XVII
P4 Pineus pinifoliae IV,XII XV
(
R1 Resselieila spp.
R2 Rhabdophaga swainei
R3 Rhyacionia buoliana IV,VI, XVI, XVII
S1 Spilonota lariciana
T1 Tetyra bipunctata
T2 Tourneyella parvicornis XII
X1 Xyela spp.
I Acephate ( orthene) 75 SP
IV Diazinon 50 EC
V Diazinon 50 WP
VII Dylox 80 SP
VIII Genite 50 EC
IX Kelthane 18.5% EC
X Kelthane 18.5 % WP
XI Lindane 20 EC
XII Malathion 50 EC
.
P6 T2 Z1 . .
02,03,P2 P3, P 4, P5,
XIII Methoxychlor 50 W
XIV Methoxychlor 24 E
XVII Sevin 50 W
Agriculture Canada. 1984 . Compendium of pest control products registered in Canada: Control of arthropods
and molluscs . Agric. Can. Pesticides Division, Plant Health and Plant Products Directorate. 1984.
Brown, N .R.; Amirault, P. A . 1985. Studies on the biology and control of cone and seed insects of selected
conifers in the Maritime Provinces. Final Report: Contract No: 08 SC.KH209-3-0137 Supply and Services.
Canada ( for Agriculture Canada) 65p.
APPENDIX X
Column
No.
It is important to record the dates of the different stages of tree development. This information can
be “tied-in” with weather data and used for operations such as frost protection ( Determine if the
trees are at a ‘frost-sensitive’ stage BEFORE an expected heavy frost and then assess if frost
protection is necessary ).
20- 23 Year
Dates at which MOST of the buds have flushed, e. g., for the spruces, bud cap has separated from
the base of the bud .
24- 27 Seed cones
32- 35 Vegetative
36- 39 Start: The first date that pollen is released when the buds / branches are shaken.
40- 43 End: Pollen shed is deemed as finished when the spent pollen cones start to dry out and little pollen
is released when the buds/ branches are shaken.
59- 80 COMMENTS
X- 4
/
X- 3
CL
co CD
CC
o NUMBER 9
M D
cf0 VEG, START END START END LEADER TREE
LENGTH D 1 AM
o M M D M D M D M D M D (cm) y in m)
I
2 3 4 5 6 7 8 9 10 I ! 12 13 14 15 16 ( 7 10 19 2C 21 22 23 24 25 2627 2829 3031 3233 34 35 363738 39 4641 42 43 44 45 46 47 46 49 50 51 52 53154 55 56 57 58 59 60 61 62 63 6465 66676869 70 71 72 73 74 75 76 77 76 7960
XII-1
APPENDIX XII
GLOSSARY
Clone: A group of genetically identical plants derived asexually from a single individual by grafting, rooting
cuttings or tissue culture techniques.
Cone: One of the reproductive structures of conifers. A female cone or seed cone bears seeds while the male
cone or pollen cone bears pollen.
Conelet: An immature female cone of conifers. A young cone from the time following pollination until it has
almost attained full size before maturity.
Contamination: The introduction of foreign matter that may alter the behavior of material under observation,
e.g ., foreign pollen in seed orchards or in single-tree pollens used for controlled pollination.
Controlled pollination: The transfer of pollen from a known tree to the female strobili of another known tree
excluding any foreign pollen.
Cross (noun): The plant resulting when two plants which are different genetically are control pollinated
[ hybrid ], ( verb ) To cross-polImate [ hydridize ].
Cutting: Detached portion of leaf , stem, or piece of root which is encouraged to form roots thus producing an
entire plant.
Embryo: The portion of the seed resulting from the union of male and female gametes which develops into a
mature plant.
Family, full- sib: The offspring of a single pair of trees, usually resulting from controlled pollination.
Family, half - sib: The offspring of a single tree ( usually female parent) having different parents of the other sex.
A half -sib family may result from open pollination or from controlled pollination using a mixture of pollen.
Family test: Evaluation of groups of open-pollinated seedlings originating from separate plus trees. The
information is used to rogue seedling seed orchards and to indicate superiority of the plus tree if so desired.
Gamete: The single male cell ( pollen ) and female cell (egg) which form the zygote that develops into the
embryo.
Gametophyte: ( in a coniferous seed) Food storage tissue contained in the seed and surrounding the embryo.
Genotype- environment interaction: The reaction of trees from different families or clones to various external
stimuli, e.g., fertilizer, climate.
Graft: The completed or successful union of a detached branch ( scion) with a rootstock.
XII- 2
Graft incompatibility: A failure of the scion and rootstock to maintain a successful union due to physiological
factors.
Microstrobilus: The reproductive structure on conifers which bears pollen i.e., the pollen cone.
Open pollination: Natural pollination effected by wind or insects and not influenced directly by man .
Phenotype: The result of the interaction between genetic make-up of an individual ( genotype) and the
environment . The tree as we see it.
Pollen cone: The male reproductive structure on conifers which bears pollen.
Pollen dilution zone: An area adjacent to or surrounding a seed orchard within which most foreign pollen will
settle out.
Pollination: The act of bringing pollen into close contact with a receptive female strobilus.
Primordium (pi. primoridia): A microscopic mound of tissue (group of cells) at its earliest stage of
development of an organ, e.g., bud primordium, leaf primordium. Two phases of development are involved.
During the initiation phase, the mound of tissue forms. The differentiation stage occurs later when the
tissue develops into a particular type of bud ( vegetative, pollen cone, or seed cone) , or it may not
differentiate and remain latent.
Progeny test: Evaluation of a parent ( or parents) from the performance of its (their ) offspring (progeny ) . The
seed from these tests is usually derived from controlled pollinations.
Seed cone: The female reproductive structure of conifers which bears seeds .
Seed orchard: An artificial population of trees, isolated to reduce influx of genetically inferior pollen and
intensively managed to produce early, frequent and abundant cone crops.
Seed orchard, clonal: A seed orchard composed of vegetatively propagated trees (grafts /rooted cuttings).
Seedlot: A group of seeds used to describe a collection from a single tree, group of trees, seed orchard block,
etc.
Self -pollination: The pollinaton of a female strobilus with pollen from the same tree or clone.
.
Strobilus: (pi. strobili): One of the reproductive structures of conifers A female cone or megastrobilus bears
seeds while the male cone or microstrobilus bears pollen.
Systemic insecticide: A chemical, when applied either externally or internally to various parts of a tree, is
absorbed and translocated to untreated plant tissue, rendering the tissue toxic to insects.
Topophysis: A physiological condition whereby a grafted scion continues to grow in a branch- like habit
instead of upright.
Anonymous. 198 . Management techniques for seed orchards. Ontario Min. Nat. Res., Tree Seed and Forest
Genetics Unit., Draft.
Snyder , E.S. 1972. Glossary for forest tree improvement workers. USDA, For. Serv., South. For. Exp, Sta., New
Orleans.
Wright, J.W. 1976. Introduction to forest genetics. Academic Press Inc., New York.