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Reptiles of Western Cape [South Africa] Related checklists:  South Africa Bibliography of the reptiles of Western Cape Taxonomic history of the reptiles of Western Cape Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989   Agama atra Southern Rock Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Tolley, Whitaker & Turner 2006   Agama hispida Southern Spiny Agama Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Cornelius, Marais & Cunningham 2012   Agama knobeli Namibian Black Agama Branch 1998  Family Chamaeleonidae (Chameleons) Genus Bradypodion (Southern African Dwarf Chameleons)   * Bradypodion atromontanum Swartberg Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch, Tolley & Tilbury 2006 Glaw 2015 Tilbury 2010   Bradypodion damaranum Knysna Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1989 Branch, Tolley & Tilbury 2006 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   Bradypodion gutturale Little Karroo Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch, Tolley & Tilbury 2006 Burger 1993 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   Bradypodion occidentale Namaqua Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch, Tolley & Tilbury 2006 Tilbury 2010 Visser 1979   * Bradypodion pumilum Western Cape Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch, Tolley & Tilbury 2006 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   Bradypodion ventrale Eastern Cape Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 [as karrooicum] Branch 1981 Branch, Tolley & Tilbury 2006 Glaw 2015 Tilbury 2010   * Bradypodion venustum Grootvadersbosch Dwarf Chameleon Tolley, Tilbury & Burger 2022 Genus Chamaeleo (Typical Chameleons)   Chamaeleo namaquensis Namaqua Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Leuteritz 2005 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Chamaesaura (African Grass Lizards)   Chamaesaura anguina Southern Highland Grass Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch 1981 Branch 1989 Branch 1998 Toit, Mouton & Flemming 2003 Genus Cordylus (Girdled Lizards)   * Cordylus aridus Dwarf Karoo Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Mouton & Wyk 1994   Cordylus cordylus Cape Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1989 Branch 1998 Burger 1993 Diedericks & Daniels 2014 Heerden, Mouton & Wyk 1992 Phelps 2009 Tolley, Whitaker & Turner 2006   Cordylus macropholis Large-scaled Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 McLachlan 1979   Cordylus mclachlani Ceres Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Cordylus minor Dwarf Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1989 Branch 1998 Burger 1993   * Cordylus niger Black Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Heerden, Mouton & Wyk 1992 Witberg 2012   * Cordylus oelofseni Porterville Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Mouton & Wyk 1990 Tolley, Whitaker & Turner 2006 Genus Hemicordylus (Western Crag Lizards)   Hemicordylus capensis Graceful Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch 1981 Branch 1989 Branch 1998 Tolley, Whitaker & Turner 2006   * Hemicordylus nebulosus Dwarf Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Mouton & Wyk 1995 Genus Karusasaurus (Karoo Girdled Lizards)   Karusasaurus polyzonus Southern Karoo Girdled Lizard Bates 1991 Branch & Bauer 1995 Branch & Braack 1989 Branch 1989 Branch 1998 Burger 1993 Tolley, Whitaker & Turner 2006 Genus Namazonurus (Namaqua Girdled Lizards)   Namazonurus peersi Garies Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Ninurta (Blue-spotted Girdled Lizard)   Ninurta coeruleopunctatus Blue-spotted Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Genus Ouroborus (Armadillo Girdled Lizard)   Ouroborus cataphractus Armadillo Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Burger 1993 Visagie, Mouton & Flemming 2002 Genus Pseudocordylus (Eastern Crag Lizards)   Pseudocordylus microlepidotus Cape Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   * Afroedura hawequensis Hawequa Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014   Afroedura karroica Karoo Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Jacobsen, Kuhn, Jackman & Bauer 2014 Genus Afrogecko (Marbled Leaf-toed Gecko)   Afrogecko porphyreus Marbled Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch & Bauer 1995 Branch 1981 Branch 1998 Branch, Bauer & Good 1995 Heinicke, Daza, Greenbaum, Jackman & Bauer 2014 Tolley, Whitaker & Turner 2006 Witberg 2012 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus angulifer Giant Ground Gecko Branch & Braack 1989 Heinz, Brennan, Jackman & Bauer 2021 Visser 1981   Chondrodactylus bibronii Tubercled Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer 1991 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Heinz, Brennan, Jackman & Bauer 2021 Tolley, Whitaker & Turner 2006 Genus Goggia (South African Dwarf Leaf-toed Geckos)   * Goggia braacki Western Cape Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch 1998 Good, Bauer & Branch 1996 Heinicke, Turk & Bauer 2017   * Goggia essexi Eastern Cape Dwarf Leaf-toed Gecko Heinicke, Turk & Bauer 2017   Goggia hewitti Swartberg Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch, Bauer & Good 1995 Heinicke, Turk & Bauer 2017   Goggia hexapora Cedarberg Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch 1998 Branch, Bauer & Good 1995 Good, Bauer & Branch 1996 Heinicke, Turk & Bauer 2017 Tolley, Whitaker & Turner 2006   * Goggia incognita Jacobsbaai Dwarf Leaf-toed Gecko Heinicke, Turk & Bauer 2017   Goggia lineata Striped Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Branch, Bauer & Good 1995 Heinicke, Turk & Bauer 2017 Witberg 2012   Goggia matzikamaensis Matzikama Dwarf Leaf-toed Gecko Heinicke, Turk & Bauer 2017   * Goggia microlepidota Small-scaled Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch 1981 Branch 1998 Branch, Bauer & Good 1995 Heinicke, Turk & Bauer 2017 McLachlan 1979 Mouton & Wyk 1981 Visser 1979 Genus Pachydactylus (Typical Thick-toed Geckos)   Pachydactylus austeni Port Nolloth Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Pachydactylus barnardi Garies Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pachydactylus capensis Common Cape Thick-toed Gecko Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998   Pachydactylus formosus Karoo Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1981 Branch 1989 Branch 1998 Lamb & Bauer 2000 Tolley, Whitaker & Turner 2006 Visser 1979   Pachydactylus geitje Ocellated Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Bauer 1991 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1989 Branch 1998 Burger 1993 Tolley, Whitaker & Turner 2006   Pachydactylus kladaroderma Thin-skinned Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch & Braack 1989 [as namaquensis] Branch 1981 [as namaquensis] Branch 1989 [as namaquensis] Branch 1998 Branch, Bauer & Good 1996 Burger 1993 [as namaquensis]   Pachydactylus labialis Western Cape Thick-toed Gecko Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Visser 1979   Pachydactylus latirostris Broad-nosed Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Visser 1979   Pachydactylus maculatus Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Visser 1979   Pachydactylus mariquensis Marico Thick-toed Gecko Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Mouton & Mostert 1984   Pachydactylus oculatus Golden Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2005 Branch & Bauer 1995 Branch & Braack 1989 Burger 1993   Pachydactylus purcelli Karoo Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer 1991 Bauer, Lamb & Branch 2006 Branch & Braack 1989 Branch 1981 Branch 1989 Burger 1993 Nicolau & Lynch 2019   Pachydactylus weberi Klipfontein Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Lamb & Branch 2006 Genus Ptenopus (African Barking Geckos)   Ptenopus garrulus Common Barking Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Genus Ramigekko (Swartberg Leaf-toed Gecko)   * Ramigekko swartbergensis Swartberg Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch 1998 Heinicke, Daza, Greenbaum, Jackman & Bauer 2014  Family Gerrhosauridae (Plated Lizards) Genus Cordylosaurus (Dwarf Plated Lizard)   Cordylosaurus subtessellatus Dwarf Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 Visser 1979   Gerrhosaurus typicus Namaqua Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Baard 1987 Cloete, Mohamed, Ponton, Tokota & Maritz 2019 Genus Tetradactylus (Plated Snake Lizards)   * Tetradactylus fitzsimonsi Cape Plated Snake Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Tetradactylus seps Short-legged Plated Snake Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch 1981 Branch 1989 Branch 1998   Tetradactylus tetradactylus Four-toed Plated Snake Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1989 Branch 1998 Burger 1993  Family Lacertidae (Typical Lizards) Genus Australolacerta (Cape Rock Lizard)   * Australolacerta australis Cape Rock Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Villiers, Baard & Branch 1983 Genus Meroles (Southwest African Desert Lizards)   Meroles ctenodactylus Giant Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981   Meroles knoxii Round-snouted Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Bauer 2004 Branch 1989 Branch 1998 Burger 1993   Meroles suborbitalis Spotted Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1989 Genus Nucras (Sandveld Lizards)   * Nucras aurantiaca Golden Sandveld Lizard Bauer, Childers, Broeckhoven & Mouton 2019   Nucras lalandii Southeastern Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1989 Branch 1998   Nucras livida Karoo Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Broadley 1972 Conradie, Busschau & Jordaan 2020   Nucras tessellata Western Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch 1981 Branch 1998 Broadley 1972 Burger 1993 Tolley, Whitaker & Turner 2006 Genus Pedioplanis (African Sand Lizards)   Pedioplanis burchelli Southern Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1989 Branch 1998 Burger 1993   Pedioplanis laticeps Cape Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Kirchhof, Penner, Roedel & Muller 2017   Pedioplanis namaquensis Namaqua Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1989   Pedioplanis pulchella Venus Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer 1991 [as lineoocellata] Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Burger 1993 Tolley, Whitaker & Turner 2006 Genus Tropidosaura (South African Mountain Lizards)   Tropidosaura gularis Cape Mountain Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bischoff 1992 Branch & Bauer 1995 Branch 1981 Branch 1998   Tropidosaura montana Common South African Mountain Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bischoff 1992 Branch & Bauer 1995 Branch 1981 Branch 1998  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   * Acontias grayi Graafwater Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley & Greer 1969   Acontias lineatus Striped Dwarf Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 [as "cf. lineatus"] Branch 1981 Branch 1998 Broadley & Greer 1969   Acontias litoralis Coastal Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley & Greer 1969   Acontias meleagris Cape Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1998 Broadley & Greer 1969 Burger 1993 Girard 2002 Lamb, Biswas & Bauer 2010 Tolley, Whitaker & Turner 2006 Witberg 2012   Acontias tristis Northern Cape Striped Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Scelotes (Dwarf Burrowing Skinks)   * Scelotes bipes Silvery Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011   Scelotes caffer Cape Three-toed Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch 1989 Branch 1998 Burger 1993   * Scelotes gronovii Cape One-toed Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011 Witberg 2012   * Scelotes kasneri Cape Two-toed Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011   * Scelotes montispectus Bloubergstrand Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Whiting & Sadlier 2003 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011   Scelotes sexlineatus Striped Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis capensis Cape Skink Bauer 1991 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Broadley 2000 Burger 1993 Phelps 2009 Tolley, Whitaker & Turner 2006 Witberg 2012   Trachylepis homalocephala Red-sided Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Branch 1989 Branch 1998 Broadley 2000 Portik, Bauer & Jackman 2010 Tolley, Whitaker & Turner 2006   Trachylepis occidentalis Western Three-striped Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Broadley 2000 Burger 1993   Trachylepis sulcata Koppie Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer 1991 Branch & Bauer 1995 Branch & Braack 1989 Branch 1989 Broadley 2000 Burger 1993 Portik, Bauer & Jackman 2010 Portik, Bauer & Jackman 2011   Trachylepis variegata Western Variegated Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Bauer 1995 Branch & Braack 1989 Branch 1981 Broadley 1975 Burger 1993 Portik, Bauer & Jackman 2010 Tolley, Whitaker & Turner 2006 Visser 1981 Genus Typhlosaurus (Blind Legless Skinks)   * Typhlosaurus caecus Yellow Blind Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Heideman, Wilson, Hendricks, Don & Moses 1998 Branch 1981 Branch 1998 Broadley 1968 Visser 1979  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1981 Branch 1989 Els 2005

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983 Burger 1993 Genus Dasypeltis (African Egg Eaters)   Dasypeltis scabra Common African Egg Eater Bates & Broadley 2018 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Broadley 1983 Burger 1993 Tolley, Whitaker & Turner 2006 Witberg 2012 Genus Dispholidus (Boomslangs)   Dispholidus typus Cape Boomslang Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Wallach 2002 Broadley 1983 Burger 1993 Visser 1979 Genus Philothamnus (African Green Snakes)   Philothamnus hoplogaster Southeastern Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Jacobsen 2006   Philothamnus occidentalis Western Natal Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Jesus, Nagy, Branch, Wink, Brehm & Harris 2009 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus beetzii Namib Tiger Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Baard 1987 Visser 1981  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps lubricus African Coral Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1989 Branch 1998 Broadley 1983 Visser 1979 Wallach, Williams & Boundy 2014 Genus Hemachatus (Ring-necked Spitting Cobra)   Hemachatus haemachates Ring-necked Spitting Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983 Smith 2011 Visser 1979 Genus Naja (Typical Cobras)   Naja nigricincta Western Barred Spitting Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1968 Broadley 1983 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja nivea Yellow Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley & Wuster 2004 Broadley 1983 Burger 1993 Dobiey & Vogel 2007  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis platurus Yellow-bellied Sea Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983  Family Lamprophiidae (House Snakes, etc.) Genus Alopecion (Spotted House Snake)   Alopecion guttatum Spotted House Snake Branch & Braack 1989 Branch 1989 Broadley 1983 Burger 1993 Tolley, Whitaker & Turner 2006 Visser 1979 Visser 1981 Genus Amplorhinus (Many-spotted Snake)   Amplorhinus multimaculatus Many-spotted Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Broadley 1983 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 [as fuliginosus] Broadley 1983 [as fuliginosus] Tolley, Whitaker & Turner 2006 [as fuliginosus] Visser 1979 [as fuliginosus] Genus Dipsina (Dwarf Beaked Snake)   Dipsina multimaculata Dwarf Beaked Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Broadley 1983 Baard 1987 Visser 1979 Genus Duberria (African Slug Eaters)   Duberria lutrix Common Slug Eater Branch & Braack 1989 Broadley 1983 Burger 1993 Genus Homoroselaps (African Harlequin Snakes)   Homoroselaps lacteus Spotted Harlequin Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1989 Broadley 1983 Burger 1993 Dobiey & Vogel 2007 Genus Lamprophis (SubSaharan House Snakes)   Lamprophis aurora Aurora House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983   Lamprophis fiskii Cape House Snake Barts, Schneider, Boone, Marais, Branch & Haacke 2012 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Baard 1987 Marais 1981   Lamprophis fuscus Yellow-bellied House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Visser 1979 Genus Lycodonomorphus (Southern African Water Snakes)   Lycodonomorphus inornatus Olive-brown House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983   Lycodonomorphus rufulus Common Brown Water Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Branch 1998 Broadley 1983 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Broadley 1983 Broadley 1996 Genus Prosymna (Shovel-snout Snakes)   Prosymna sundevallii Southern African Shovel-snout Branch & Braack 1989 Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1983 Visser 1979 Visser 1981 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis crucifer Cross-marked Grass Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1989 Broadley 1983 Broadley 2002 Burger 1993   * Psammophis leightoni Cape Sand Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002   Psammophis namibensis Namib Sand Snake Broadley 1977 Broadley 1983 Tolley, Whitaker & Turner 2006   Psammophis notostictus Karoo Sand Snake Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1977 Broadley 1983 Burger 1993 Tolley, Whitaker & Turner 2006 Genus Psammophylax (Skaapstekers)   Psammophylax rhombeatus Spotted Skaapsteker Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983 Burger 1993 Keates, Conradie, Greenbaum & Edwards 2019 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops nigricans South African Black Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Broadley 1999 Broadley 1983 Genus Namibiana (Southwest African Thread Snakes)   Namibiana gracilior Slender Thread Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Broadley 1999 Broadley 1983 Burger 1993 McDiarmid, Campbell & Toure 1999  Family Typhlopidae (Typical Blind Snakes) Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops lalandei Cape Blind Snake Branch & Bauer 2004 Branch & Braack 1989 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Burger 1993  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Broadley 1983 Burger 1993 Phelps 2009 Tolley, Whitaker & Turner 2006 Witberg 2012   * Bitis armata Southern Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Branch 1999 Dobiey & Vogel 2007   Bitis atropos Cape Mountain Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Dobiey & Vogel 2007 McDiarmid, Campbell & Toure 1999 Visser 1979   Bitis caudalis Horned Adder Branch & Braack 1989 Broadley 1983 Dobiey & Vogel 2007 Phelps & Els 2006   Bitis cornuta Many-horned Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Branch 1999 Broadley 1983 Visser 1979   Bitis rubida South African Red Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1997 Branch 1998 Branch 1999 Broadley 1983 [as inornata] Dobiey & Vogel 2007   Bitis schneideri Namaqua Dwarf Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Causus (Night Adders)   Causus rhombeatus Rhombic Night Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Dobiey & Vogel 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Branch 1998  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa galeata South African Helmeted Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as subrufa] Branch & Braack 1989 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Stuart & Meakin 1983 [as P. subrufa] Tolley, Whitaker & Turner 2006 [as subrufa]  Family Testudinidae (Tortoises) Genus Chersina (Angulate Tortoise)   Chersina angulata Angulate Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Burger 1993 Hofmeyr 2009 Phelps 2009 Tolley, Whitaker & Turner 2006 Genus Chersobius (Five-toed Dwarf Tortoises)   Chersobius boulengeri Karroo Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998   Chersobius signatus Speckled Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Boycott 1986 Branch 1998 Daniels, Hofmeyr, Henen & Baard 2010 Genus Homopus (Four-toed Dwarf Tortoises)   Homopus areolatus Common Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Burger 1993 Fritz & Havas 2006 Tolley, Whitaker & Turner 2006   Homopus femoralis Greater Padloper Branch & Braack 1989 Branch 1998 Burger 1993 Fritz & Havas 2006 Genus Psammobates (South African Star Tortoises)   Psammobates geometricus Geometric Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Baard 1992 Fritz & Havas 2006   Psammobates tentorius African Tent Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Branch 1998 Burger 1993 Fritz & Havas 2006 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Braack 1989 Burger 1993