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Invertebrates: Separated the ctenophore from the cnidarian paragraph, as they refer to different phyla of animals
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{{Short description|Varied organ found in many animals and used for palpation and manipulation}}
{{about||the Italian-American horror film|Tentacles (film)|the album by Merzbow|Tentacle (album)}}
{{About|tentacles in zoology}}
{{Use dmy dates|date=March 2017}}
[[Image:cuttlefish.png|thumb|upright|[[Cuttlefish]] with 2two tentacles and 8eight arms]]
 
In [[zoology]], a '''tentacle''' is a flexible, mobile, and elongated [[organ (anatomy)|organ]] present in some species of [[animal]]s, most of them [[invertebrate]]s. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work mainly like [[muscular hydrostat]]s. Most forms of tentacles are used for grasping and feeding. Many are [[Sensory system|sensory organorgans]]s, variously receptive to [[somatosensory system|touch]], [[visual perception|vision]], or to the [[Chemoreceptor|smell or taste]] of particular [[Chemoreceptor|foods or threats]]. Examples of such tentacles are the tentacles or "eye stalks"[[eyestalk]]s of various kinds of [[snail]]s. Some kinds of tentacles have both sensory and manipulatory functions. (im a barbie girl in the barbie world)
[[Image:cuttlefish.png|thumb|upright|[[Cuttlefish]] with 2 tentacles and 8 arms]]
 
A tentacle is similar to a [[cirrus (biology)|cirrus]], but a cirrus is an organ that usually lacks the tentacle's strength, size, flexibility, or sensitivity. A [[nautilus]] has cirri, but a [[squid]] has tentacles.
In [[zoology]] a '''tentacle''' is a flexible, mobile, elongated [[organ (anatomy)|organ]] present in some species of [[animal]]s, most of them [[invertebrate]]s. In animal anatomy tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work mainly like [[muscular hydrostat]]s. Most forms of tentacles are used for grasping and feeding. Many are [[Sensory system|sensory organ]]s, variously receptive to [[somatosensory system|touch]], [[vision]], or to the smell or taste of particular [[Chemoreceptor|foods or threats]]. Examples of such tentacles are the tentacles or "eye stalks" of various kinds of [[snail]]s. Some kinds of tentacles have both sensory and manipulatory functions. (im a barbie girl in the barbie world)
 
===Invertebrates===
The word '''tentillum''' literally means "little tentacle". However, irrespective of size, it usually refers to a side branch of a larger tentacle. In some cases such tentilla are specialised for particular functions; for example, in the [[Cnidaria]] tentilla usually bear [[cnidocyte]]s,<ref>[http://species-identification.org/species.php?species_group=zsao&selected=definitie&menuentry=woordenlijst&record=tentilla Marine Species Identification Portal : Zooplankton of the South Atlantic Ocean : Glossary : tentilla]. Species-identification.org. Retrieved on 2013-05-02.</ref> whereas in the [[Ctenophora]] they usually bear [[collocyte]]s.<ref name="CNH1">Harmer, Sir Sidney Frederic; Shipley, Arthur Everett et al. (1906) ''The Cambridge natural history Volume 1, Protozoa, Porifera, Coelenterata, Ctenophora, Echinodermata''. Macmillan Company.</ref><ref>{{cite journal|author=Mackie G.O., Mills C.E., Singla C.L.|title= Structure and function of the prehensile tentilla of ''Euplokamis'' (Ctenophora, Cydippida)|doi=10.1007/BF00312216|url=http://faculty.washington.edu/cemills/EuplokamisTentacle1988.pdf|year=1988|journal=Zoomorphology|volume=107|issue=6|pages=319}}</ref>
 
==== Molluscs= ===
In botany "tentacle" refers to the glandular hairs on the leaves of some species of [[carnivorous plant|insectivorous plants]], such as [[Drosera]]. This usage is not to be confused with organs such as the [[tendril]]s of climbing plants.
 
==Tentacles in animals==
===Invertebrates===
 
====Molluscs====
{{See also|Cephalopod limb}}
[[Image:Snail-front-0A.jpg|thumb|right|Front view of land snail showing upper and lower sets of tentacles]]
Many molluscs have tentacles of one form or another. The most familiar are those of the [[Pulmonata|Pulmonate land snails]], which usually have two sets of tentacles on the head: when extended the upper pair have eyes at their tips; the lower pair are [[chemoreceptor]]s. Both pairs are fully retractable [[muscular hydrostat]]s, but they are not used for manipulation or prey capture.
[[Image:White abalone Haliotis sorenseni.jpg|thumb|350 px|Abalone showing pallial tentacles]]
Many molluscs have tentacles of one form or another. The most familiar are those of the [[Pulmonata|Pulmonatepulmonate land snails]], which usually have two sets of tentacles on the head: when extended the upper pair have eyes at their tips; the lower pair are [[chemoreceptor]]s. Both pairs are fully retractable [[muscular hydrostat]]s, but they are not used for manipulation or prey capture. Molluscs have one pair of tentacles close to their mouths that hold close to their captured prey before they can consume it.<ref name="FiCaDB">{{cite journal |last1=Kier |first1=William M. |date=February 2016 |title=The Musculature of Coleoid Cephalopod Arms and Tentacles |journal=Frontiers in Cell and Developmental Biology |volume=4 |page=10 |doi=10.3389/fcell.2016.00010|doi-access=free |pmid=26925401 |pmc=4757648 }}</ref>
 
Some marine snails such as [[abalone]] and top snails, [[Trochidae]], have numerous small tentacles around the edge of the [[mantle (mollusc)|mantle]]. These are known as ''pallial tentacles''.<ref name=bou/>
 
Among [[Cephalopodacephalopod]] such as octopus ands, squid have the most spectacular tentacles. They take the form of highly mobile [[muscular hydrostat]]shydrostats with various appendages such as suction disks and sometimes thorny hooks. Up to the early twentieth century they"tentacles" were interchangeably called "arms" or "tentacles".<ref name= "CNH3">{{cite book |last1=Cooke, |first1= A. H.; |last2=Shipley, |first2=Arthur Everett et alia:|date=1895 |title=The Cambridge naturalNatural history VolumeHistory |volume=34,: Molluscs, Trilobites, Brachiopods etc. |location=London, England, U.K. |publisher= Macmillan Company}}{{page 1895needed|date=October 2023}}</ref> TheThese moderntentacles conventionare however,made isof to speakstalks of appendagesaxial as[[ventral "tentacles"nerve whencord|nerve they have relatively thin "[[pedunclecord]]s" orthat "stalks"are withcovered "clubs"by atcircular theirtransverse tips.muscle Intissue contrastthat thecontract conventionin refersresponse to the relatively shorter, [[sessile]] appendages as "arms"stimuli. ByThere thisis definitiona the eight appendageslayer of octopuses,helical thoughmuscle quitethat long,helps counteach astentacle arms,to becausetwist theyor are sessile. In terms of this article however, both types of organs count as tentacles becauseturn in relevantany waysdirection theywhere arethe structurallyprey and functionallyis similarsensed.<ref name=bou>{{cite web| author= Boumis R | title= Animals With Tentacles |url= http:"FiCaDB"//animals.pawnation.com/animals-tentacles-2495.html |date= 2013|work= Pawnation |publisher= AOL Inc. |accessdate=2013-06-08}}</ref>
 
The modern convention, however, is to speak of appendages as "tentacles" when they have relatively thin "[[peduncle (anatomy)|peduncles]]" or "stalks" with "clubs" at their tips. In contrast the convention refers to the relatively shorter appendages as "arms". By this definition the eight appendages of octopuses, though quite long, count as arms.<ref name=bou>{{cite web |author= Boumis R |title= Animals With Tentacles |url= http://animals.pawnation.com/animals-tentacles-2495.html |year= 2013 |work= Pawnation |publisher= AOL Inc. |access-date= 2013-06-08 |url-status= live |archive-url= https://web.archive.org/web/20131203020936/http://animals.pawnation.com/animals-tentacles-2495.html |archive-date= 3 December 2013 |df= dmy-all }}</ref> While arms are distinct from tentacles (a definition specific to the limb featuring peduncles), arms do fall within the general definition of "tentacle" as "a flexible, mobile, and elongated organ" and "tentacle" could be used as an [[umbrella term]].
The tentacles of the [[giant squid]] and [[colossal squid]] have powerful suckers and pointed teeth at the ends. The teeth of the giant squid resemble [[crown cork|bottle caps]], and function like tiny, circular saws; while the tentacles of the colossal squid wield two long rows of swiveling, tri-pointed hooks.
 
The tentacles of the [[giant squid]] and [[colossal squid]] have powerful suckers and pointed teeth at the ends. The teeth of the giant squid resemble [[crown cork|bottle caps]], and function like tiny, circular[[hole saws;saw]]s, while the tentacles of the colossal squid wield two long rows of swiveling, tri-pointed hooks.
[[Image:Snail-front-0A.jpg|thumb|right|Front view of land snail showing upper and lower sets of tentacles]]
[[Image:White abalone Haliotis sorenseni.jpg|thumb|350 px|Abalone showing pallial tentacles]]
 
====Cnidarians====
[[Cnidarian]]s, such as [[jellyfish]], [[sea anemone]]s, ''[[Hydra (genus)|Hydra]]'' and [[coral]] have numerous hair-like tentacles. Cnidarians have huge numbers of [[cnidocyte]]s on their tentacles. In [[jellyfish|medusoid form]], the body floats on water so that the tentacles hang down in a ring around the mouth. In [[polyp (zoology)|polyp]] form, such as sea anemone and coral, the body is below with the tentacles pointed upwards.
 
The tentacles of the [[Lionlion's mane jellyfish]] may be up to 120 feet ({{convert|37 meters)|m|ft|abbr=on}} long. They are hollow and are arranged in 8 groups of between 70 and 150. The longer tentacles are equipped with ctenophorescnidocytes whose venom paralyses and kills prey. The smaller tentacles guide food into the mouth.<ref>{{cite web | author= Kosner AW | title= Lion's Mane Jellyfish Image: This Is (Literally) How Things Blow Up On The Internet! | url= httphttps://www.forbes.com/sites/anthonykosner/2012/07/10/lions-mane-jellyfish-image-this-is-literally-how-things-blow-up-on-the-internet/2/ | date= 10 July 2012 | work= [[Forbes]] |publisher access-date= Forbes2013-06-08 | url-status= live | archive-url= https://web.archive.org/web/20130625154715/http://www.forbes.com/sites/anthonykosner/2012/07/10/lions-mane-jellyfish-image-this-is-literally-how-things-blow-up-on-the-internet/2/ LLC™|accessdate archive-date= 25 June 2013 | df= dmy-06-08all }}</ref><ref>{{cite web| author= Wild Facts | title= Wild Fact #419 – One Large Jelly – Lion’sLion's Mane Jellyfish|url=http://www.wild-facts.com/2011/wild-fact-419-one-large-jelly-lions-mane-jellyfish/|date= 29 November 2011 |work= wild-facts.com|publisher= |accessdateaccess-date=2013-06-08|url-status=live|archive-url=https://web.archive.org/web/20121219091103/http://www.wild-facts.com/2011/wild-fact-419-one-large-jelly-lions-mane-jellyfish/|archive-date=19 December 2012|df=dmy-all}}</ref>
====Cnidarians====
[[Cnidarian]]s, such as [[jellyfish]], [[sea anemone]]s, ''[[Hydra]]'' and [[coral]] have numerous hair-like tentacles. Cnidarians have huge numbers of [[cnidocyte]]s on their tentacles. In [[jellyfish|medusoid form]], the body floats on water so that the tentacles hang down in a ring around the mouth. In [[polyp]] form, such as sea anemone and coral, the body is below with the tentacles pointed upwards.
Many species of the jellyfish-like [[ctenophore]]s have two tentacles, while some have none. Their tentacles have adhesive structures called ''[[colloblast]]s'' or ''lasso cells''. The colloblasts burst open when prey comes in contact with the tentacle, releasing sticky threads that secure the food.<ref>{{cite web| author= Bird J | title= CNIDARIANS: SIMPLE ANIMALS WITH A STING! |url= http://www.oceanicresearch.org/education/wonders/cnidarian.html|date= 5 June 2007|work= oceanicresearch.org |publisher= Oceanic Research Group |accessdate=2013-06-08}}</ref>
 
===Ctenophores===
The tentacles of the [[Lion's mane jellyfish]] may be up to 120 feet (37 meters) long. They are hollow and are arranged in 8 groups of between 70 and 150. The longer tentacles are equipped with ctenophores whose venom paralyses and kills prey. The smaller tentacles guide food into the mouth.<ref>{{cite web| author= Kosner AW | title= Lion's Mane Jellyfish Image: This Is (Literally) How Things Blow Up On The Internet! |url= http://www.forbes.com/sites/anthonykosner/2012/07/10/lions-mane-jellyfish-image-this-is-literally-how-things-blow-up-on-the-internet/2/|date= 10 July 2012 |work= Forbes |publisher= Forbes.com LLC™|accessdate=2013-06-08}}</ref><ref>{{cite web| author= Wild Facts | title= Wild Fact #419 – One Large Jelly – Lion’s Mane Jellyfish|url=http://www.wild-facts.com/2011/wild-fact-419-one-large-jelly-lions-mane-jellyfish/|date= 29 November 2011 |work= wild-facts.com|publisher= |accessdate=2013-06-08}}</ref>
Many species of the jellyfish-like [[ctenophore]]s have two tentacles, while some have none. Their tentacles have adhesive structures called ''[[colloblast]]s'' or ''lasso cells''. The colloblasts burst open when prey comes in contact with the tentacle, releasing sticky threads that secure the food.<ref>{{cite web | author= Bird J | title= CNIDARIANS: SIMPLE ANIMALS WITH A STING! | url= http://www.oceanicresearch.org/education/wonders/cnidarian.html | date= 5 June 2007 | work= oceanicresearch.org | publisher= Oceanic Research Group |accessdate access-date= 2013-06-08 | url-status= live | archive-url= https://web.archive.org/web/20130707024559/http://www.oceanicresearch.org/education/wonders/cnidarian.html | archive-date= 7 July 2013 | df= dmy-all }}</ref>
 
====Bryozoan=Bryozoa===
 
[[Bryozoa]] (moss animals) are tiny creatures with tentacles around their mouths. The tentacles are almost cylindrical and have bands of cilia which create a water current towards the mouth. The animal extracts edible material from the flow of water.<ref>{{citeCite book web| author= Claus N | title= Bryozoa (Ectoprocta: ‘Moss’'Moss' Animals) | url= http://www.els.net/WileyCDA/ElsArticle/refId-a0001613.html | date= May 2013 | work= els.net | publisher= eLS. John Wiley & Sons Ltd | doi= 10.1002/9780470015902.a0001613.pub2 |accessdate access-date= 2013-06-08 | url-status= live | archive-url= https://web.archive.org/web/20130605003827/http://www.els.net/WileyCDA/ElsArticle/refId-a0001613.html | archive-date= 5 June 2013 | df= dmy-all | isbn= 978-0470016176 }}</ref>
 
===Trypanorhynch cestodes===
 
{{multiple image
| align = right
| direction = horizontal
| width = 130
| image1 = Parasite140092-fig3 - FIG11 Nybelinia basimegacantha body.png
| alt1 =
| caption1 = A larva of [[trypanorhynch]] cestode (only two tentacles shown). Scale-bar: 0.1 mm
| image2 = FIG11 Nybelinia basimegacantha Tentacle.png
| alt2 =
| caption2 = Detail of one tentacle with its spines. Scale-bar: 0.01 mm.
}}
 
[[Trypanorhynch]] [[cestode]]s are parasitic in fish. Their [[scolex]] shows four tentacles which are covered by spines. These tentacles help the adult cestode to attach to the intestine of the shark or ray that they parasitize. The same tentacles are also present in the larvae.<ref name="BeveridgeBray2014">{{cite journal|last1=Beveridge|first1=Ian|last2=Bray|first2=Rodney A.|last3=Cribb|first3=Thomas H.|last4=Justine|first4=Jean-Lou|title=Diversity of trypanorhynch metacestodes in teleost fishes from coral reefs off eastern Australia and New Caledonia|journal=Parasite|volume=21|year=2014|pages=60|issn=1776-1042|doi=10.1051/parasite/2014060|pmid=25402635|pmc=4234045|url=https://www.parasite-journal.org/articles/parasite/full_html/2014/01/parasite140092/parasite140092.html|url-status=live|archive-url=https://web.archive.org/web/20180109144048/https://www.parasite-journal.org/articles/parasite/full_html/2014/01/parasite140092/parasite140092.html|archive-date=9 January 2018|df=dmy-all}} {{open access}}</ref>
 
==Vertebrates==
 
===Amphibians===
Line 40 ⟶ 58:
 
===Mammals===
The [[star-nosed mole]], ''Condylura cristata'', of [[North America]], has 22 short but conspicuous tentacles around its nose. They are mobile and extremely sensitive, helping the animal to find its way about the burrow and detect prey. They are about 1-4 1–4&nbsp;mm long and hold about 25,000 [[Somatosensory system|touch receptors]] called [[Eimer's organ]]s, perhaps giving this mole the most delicate sense of touch among mammals.<ref name=bou/>
 
{{anchor|tentilla}}
 
==Tentillum==
==Tentacles in plants==
[[File:Ctenophore.jpg|thumb|Deep-sea ctenophore trailing tentacles studded with tentilla]]
[[Image:Drosera capensis bend.JPG|thumb|upright|Leaf and tentacle movement on ''[[Drosera capensis]]'']]
 
The word '''tentillum''' ({{plural form}}: '''tentilla''') literally means "little tentacle". However, irrespective of size, it usually refers to a side branch of a larger tentacle. In some cases, such tentilla are specialised for particular functions; for example, in the [[Cnidaria]] tentilla usually bear [[cnidocyte]]s,<ref>[http://species-identification.org/species.php?species_group=zsao&selected=definitie&menuentry=woordenlijst&record=tentilla Marine Species Identification Portal : Zooplankton of the South Atlantic Ocean : Glossary : tentilla] {{webarchive|url=https://web.archive.org/web/20131202231841/http://species-identification.org/species.php?species_group=zsao&selected=definitie&menuentry=woordenlijst&record=tentilla |date=2 December 2013 }}. Species-identification.org. Retrieved on 2013-05-02.</ref> whereas in the [[Ctenophora]] they usually bearhave [[collocyte]]s.<ref name="CNH1">Harmer, Sir Sidney Frederic; Shipley, Arthur Everett et al. (1906) ''The Cambridge naturalNatural historyHistory Volume 1, Protozoa, Porifera, Coelenterata, Ctenophora, Echinodermata''. Macmillan Company.</ref><ref>{{cite journal|authorauthor1=Mackie G.O., |author2=Mills C.E., |author3=Singla C.L.|title= Structure and function of the prehensile tentilla of ''Euplokamis'' (Ctenophora, Cydippida)|doi=10.1007/BF00312216|url=http://faculty.washington.edu/cemills/EuplokamisTentacle1988.pdf|year=1988|journal=Zoomorphology|volume=107|issue=6|pages=319|s2cid=317017|url-status=live|archive-url=https://web.archive.org/web/20160303233312/http://faculty.washington.edu/cemills/EuplokamisTentacle1988.pdf|archive-date=3 March 2016|df=dmy-all}}</ref> [[Siphonophorae|Siphonophores]] are an example of Cnidaria that use tentilla.
In [[carnivorous plant]]s, like [[Drosera|sundew]], the so-called tentacles are the stalked glands of the upper surface of the [[leaf|leaves]]. They are hairlike projections with a drop of nectar-like glue which attract insects. Unlike a usual plant hair which is of epidermal origin, the tentacle is highly complex and includes all the tissue types present in the leaf. It consists of a tall, tapering stalk of multi-cellular structure, topped with two layers of glandular cells. They are highly touch-sensitive. When an insect or small animal is captured, the tentacles bend inward and the leaf rolls together as shown in the picture. The glandular cells then secrete [[enzyme]]s to dissolve and digest the insect.<ref>{{cite web| author= Honda M | title= Carnivorous Plants / Insectivorous Plants in the Wilderness: Sundews |url= http://www.honda-e.com/IPW_3_Description/TX-3Sundews.htm|date= |work= Carnivorous Plants Web Site |publisher= |accessdate=2013-06-08}}</ref><ref>{{cite journal| author= Poppinga S, Hartmeyer SRH, Seidel R, Masselter T, Hartmeyer I, Speck T |title= Catapulting Tentacles in a Sticky Carnivorous Plant |year = 2013 |url= | journal= PLoS ONE | volume= 7 | issue= 9 |page = e45735 |pmid=23049849 |pmc= PMC3458893|doi= 10.1371/journal.pone.0045735}}</ref>
 
==References==
{{reflistReflist}}
 
==External Linklinks==
{{Commons category|Tentacles}}
*[http://animals.about.com/od/t/g/tentacle.htm Tentacle] {{Webarchive|url=https://web.archive.org/web/20130514203145/http://animals.about.com/od/t/g/tentacle.htm |date=14 May 2013 }} at [[About.com]]
*[http://www.britannica.com/EBchecked/topic/587589/tentacle EnclopaediaTentacle] at the ''[[Encyclopædia Britannica]]''
*{{wiktionary-inline}}
 
{{Authority control}}
*[http://animals.about.com/od/t/g/tentacle.htm About.com]
*[http://www.aboutsociology.com/sociology/Tentacle About Sociology]
*[http://www.britannica.com/EBchecked/topic/587589/tentacle Enclopaedia Britannica]
*[http://userwww.sfsu.edu/biol240/labs/lab_18molluscs/pages/cephalopods.html Cephalopods]
 
[[Category:Animal anatomy]]
[[Category:Animal morphology]]
[[Category:Plant morphology]]
[[Category:Carnivorous plants]]