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The cast of the 1980s space-adventure series ''Galaxy Quest'' attend [[fan convention]]s and make trivial promotional appearances. Though the series' former star, Jason Nesmith, thrives on the attention, his co-stars—Gwen, Alexander, Fred, and Tommy—resent him and the states of their careers. At a convention, a group calling themselves Thermians approaches Jason for help. Thinking they want him for a promotional appearance, he agrees. The next morning, when they pick him up, Jason is hung over and does not grasp that the Thermians are aliens who have transported him to a working re-creation of the ''Galaxy Quest'' starship, the NSEA ''Protector''. Jason believes he is on a set, and performs in character as he confronts the Thermians' enemy, Sarris, who demands the "Omega 13", a secret super weapon with unknown capabilities mentioned in the show's finale. Giving perfunctory orders, Jason manages to temporarily defeat Sarris.
After the grateful Thermians transport him back to Earth, Jason realizes the experience was real and attempts to convince the other cast members. When the Thermian Laliari seeks Jason's help again, the cast joins him, along with the convention [[Master of ceremonies|emcee]], Guy, who had played an
Sarris returns and demands the "Omega 13" device. He attacks the ''Protector'' again, and the ship barely escapes through a magnetic minefield. However, the ship's power source, a [[beryllium]] sphere, is severely damaged. The humans travel to a nearby planet and take a replacement sphere from ferocious, childlike alien miners. On their return to the ''Protector'', they discover that Sarris has seized the ship. After Jason confesses they are just actors, Sarris forces him to explain the truth to the disillusioned Thermian leader, Mathesar. Sarris activates the ''Protector'''s [[self-destruct]] mechanism and returns to his ship.
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