(usuallynowin the plural) One of the long, narrow feathers on the neck of birds, most noticeable on the rooster. [from 15th c.]
(fishing) A feather used to make a fishing lure or a fishing lure incorporating a feather. [from 17th c.]
(usuallynowin the plural) By extension (because the hackles of a rooster are lifted when it is angry), the hair on the nape of the neck in dogs and other animals; also used figuratively for humans. [from 19th c.]
When the dog got angry, his hackles rose and he growled.
1976, Richard Dawkins, The Selfish Gene, Kindle edition, OUP Oxford, published 2016, page 101:
Suppose it happened to be the case that the majority of individuals raised their hackles only when they were truly intending to go on for a very long time in the war of attrition. The obvious counterploy would evolve: individuals would give up immediately when an opponent raised his hackles.
A type of jagged crack extending inwards from the broken surface of a fractured material.
A plate with rows of pointed needles used to blend or straighten hair. [from 20th c.]
1949, Eric Philbrook Kelly, The Amazing Journey of David Ingram, page 11:
"COME ALONG YE GRASS-COMBERS, SHOW some hackle," David Ingram, striding ahead, turned back and called.
Usage notes
In everyday speech, primarily used in phrase to raise someone's hackles(“to make one angry”), as in “It raises my hackles when you take that condescending tone”.